Cu/Zn superoxide dismutase homologs participate in Nicotiana benthamiana antiviral responses

Superoxide dismutases (SODs) serve as the first line of defense against reactive oxygen species. Copper-zinc superoxide dismutase (Cu/Zn-SOD) is an enzyme whose activity depends on copper availability. Cu/Zn-SOD-1 induction is shown to be involved in the antioxidative and antiviral activity of acety...

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Main Authors: Haijuan Wang, Jidan Zhang, Zhuo Meng, Zhenqi Sun, Dongyang Liu, Bin Li, Fangfang Yan, Chongyi Jia, Hongyou Zhou, Mingmin Zhao
Format: Article
Language:English
Published: Frontiers Media S.A. 2025-07-01
Series:Frontiers in Microbiology
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Online Access:https://www.frontiersin.org/articles/10.3389/fmicb.2025.1561731/full
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Summary:Superoxide dismutases (SODs) serve as the first line of defense against reactive oxygen species. Copper-zinc superoxide dismutase (Cu/Zn-SOD) is an enzyme whose activity depends on copper availability. Cu/Zn-SOD-1 induction is shown to be involved in the antioxidative and antiviral activity of acetylsalicylic acid in hepatitis C virus (HCV)-expressing cells. Here, RNA sequencing (RNA-seq) analysis identified SOD homologs (three NbCu/Zn-SOD, four NbFe-SOD, and two NbMn-SOD) that were differentially expressed in Nicotiana benthamiana during tobacco vein mottling virus (TVMV) infection. NbCu/Zn-SOD-1 was cloned from N. benthamiana and subsequently characterized. Encoding sequence and structural analyses of the NbCu/Zn-SOD-1 protein confirmed a conserved SOD enzyme domain, GFHLHEfGDtT, indicating that it is SOD-dependent and phylogenetically related to Cu/Zn-SOD in Nicotiana tabacum (XP 016486719.1). NbCu/Zn-SOD-1 was primarily localized in the cytoplasm. The transient expression of NbCu/Zn-SOD-1 led to a reduced accumulation of TVMV and PVY-Rosea1 (PVY-Ros1). In summary, our results suggest that NbCu/Zn-SOD-1 homologs participate in plant antiviral responses.
ISSN:1664-302X