Characterization of metabolically quiescent Leishmania parasites in murine lesions using heavy water labeling.

Information on the growth rate and metabolism of microbial pathogens that cause long-term chronic infections is limited, reflecting the absence of suitable tools for measuring these parameters in vivo. Here, we have measured the replication and physiological state of Leishmania mexicana parasites in...

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Main Authors: Joachim Kloehn, Eleanor C Saunders, Sean O'Callaghan, Michael J Dagley, Malcolm J McConville
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2015-02-01
Series:PLoS Pathogens
Online Access:https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1004683&type=printable
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author Joachim Kloehn
Eleanor C Saunders
Sean O'Callaghan
Michael J Dagley
Malcolm J McConville
author_facet Joachim Kloehn
Eleanor C Saunders
Sean O'Callaghan
Michael J Dagley
Malcolm J McConville
author_sort Joachim Kloehn
collection DOAJ
description Information on the growth rate and metabolism of microbial pathogens that cause long-term chronic infections is limited, reflecting the absence of suitable tools for measuring these parameters in vivo. Here, we have measured the replication and physiological state of Leishmania mexicana parasites in murine inflammatory lesions using 2H2O labeling. Infected BALB/c mice were labeled with 2H2O for up to 4 months, and the turnover of parasite DNA, RNA, protein and membrane lipids estimated from the rate of deuterium enrichment in constituent pentose sugars, amino acids, and fatty acids, respectively. We show that the replication rate of parasite stages in these tissues is very slow (doubling time of ~12 days), but remarkably constant throughout lesion development. Lesion parasites also exhibit markedly lower rates of RNA synthesis, protein turnover and membrane lipid synthesis than parasite stages isolated from ex vivo infected macrophages or cultured in vitro, suggesting that formation of lesions induces parasites to enter a semi-quiescent physiological state. Significantly, the determined parasite growth rate accounts for the overall increase in parasite burden indicating that parasite death and turnover of infected host cells in these lesions is minimal. We propose that the Leishmania response to lesion formation is an important adaptive strategy that minimizes macrophage activation, providing a permissive environment that supports progressive expansion of parasite burden. This labeling approach can be used to measure the dynamics of other host-microbe interactions in situ.
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spelling doaj-art-ac267e0f616f4038bcbfe85880fcc2ec2025-08-20T02:15:13ZengPublic Library of Science (PLoS)PLoS Pathogens1553-73661553-73742015-02-01112e100468310.1371/journal.ppat.1004683Characterization of metabolically quiescent Leishmania parasites in murine lesions using heavy water labeling.Joachim KloehnEleanor C SaundersSean O'CallaghanMichael J DagleyMalcolm J McConvilleInformation on the growth rate and metabolism of microbial pathogens that cause long-term chronic infections is limited, reflecting the absence of suitable tools for measuring these parameters in vivo. Here, we have measured the replication and physiological state of Leishmania mexicana parasites in murine inflammatory lesions using 2H2O labeling. Infected BALB/c mice were labeled with 2H2O for up to 4 months, and the turnover of parasite DNA, RNA, protein and membrane lipids estimated from the rate of deuterium enrichment in constituent pentose sugars, amino acids, and fatty acids, respectively. We show that the replication rate of parasite stages in these tissues is very slow (doubling time of ~12 days), but remarkably constant throughout lesion development. Lesion parasites also exhibit markedly lower rates of RNA synthesis, protein turnover and membrane lipid synthesis than parasite stages isolated from ex vivo infected macrophages or cultured in vitro, suggesting that formation of lesions induces parasites to enter a semi-quiescent physiological state. Significantly, the determined parasite growth rate accounts for the overall increase in parasite burden indicating that parasite death and turnover of infected host cells in these lesions is minimal. We propose that the Leishmania response to lesion formation is an important adaptive strategy that minimizes macrophage activation, providing a permissive environment that supports progressive expansion of parasite burden. This labeling approach can be used to measure the dynamics of other host-microbe interactions in situ.https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1004683&type=printable
spellingShingle Joachim Kloehn
Eleanor C Saunders
Sean O'Callaghan
Michael J Dagley
Malcolm J McConville
Characterization of metabolically quiescent Leishmania parasites in murine lesions using heavy water labeling.
PLoS Pathogens
title Characterization of metabolically quiescent Leishmania parasites in murine lesions using heavy water labeling.
title_full Characterization of metabolically quiescent Leishmania parasites in murine lesions using heavy water labeling.
title_fullStr Characterization of metabolically quiescent Leishmania parasites in murine lesions using heavy water labeling.
title_full_unstemmed Characterization of metabolically quiescent Leishmania parasites in murine lesions using heavy water labeling.
title_short Characterization of metabolically quiescent Leishmania parasites in murine lesions using heavy water labeling.
title_sort characterization of metabolically quiescent leishmania parasites in murine lesions using heavy water labeling
url https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1004683&type=printable
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