Transcriptome and co-expression network analysis reveal the genetic basis of cell wall components in maize stalks

Abstract Cellulose (CEL), hemicellulose (HCE), and lignin (LIG) are crucial constituents of the cell wall in maize stalks, significantly influencing stalk resistance to lodging and external stresses. The proportion of these contents also determines the application of maize stalks in animal feed, ene...

Full description

Saved in:
Bibliographic Details
Main Authors: Yao He, Zhike Deng, Shijiang He, Zhaojie Qi, Hengshuo Chang, Peng Liu, Zhong Chen, Chaoying Zou, Yaou Shen, Langlang Ma
Format: Article
Language:English
Published: BMC 2025-07-01
Series:BMC Genomics
Subjects:
Online Access:https://doi.org/10.1186/s12864-025-11816-2
Tags: Add Tag
No Tags, Be the first to tag this record!
_version_ 1849403231831064576
author Yao He
Zhike Deng
Shijiang He
Zhaojie Qi
Hengshuo Chang
Peng Liu
Zhong Chen
Chaoying Zou
Yaou Shen
Langlang Ma
author_facet Yao He
Zhike Deng
Shijiang He
Zhaojie Qi
Hengshuo Chang
Peng Liu
Zhong Chen
Chaoying Zou
Yaou Shen
Langlang Ma
author_sort Yao He
collection DOAJ
description Abstract Cellulose (CEL), hemicellulose (HCE), and lignin (LIG) are crucial constituents of the cell wall in maize stalks, significantly influencing stalk resistance to lodging and external stresses. The proportion of these contents also determines the application of maize stalks in animal feed, energy production, and other fields. Herein, two maize inbred lines with contrasting cell wall component contents in the stalk, MO190 (high CEL, HCE, and LIG contents) and MO194 (low CEL, HCE, and LIG contents), were subjected to analysis of transcriptomes at the seedling, jointing, silking, and grain stages. By comparing the differentially expressed genes (DEGs) during stalk development between the two lines, 11,546 genes were identified as the specific DEGs in each line. Using traits-associated WGCNA (weight gene co-expression network analysis), we identified a hub co-expression module and 26 key genes regulating the CEL, HCE, and LIG contents in the stalk. Based on the eigengene-based connectivity value in the module, the four genes Zm00001d019317, Zm00001d027313, Zm00001d043965, and Zm00001d019149 were confirmed as the hub genes affecting these contents. Among these, Zm00001d019317 that encodes CEL synthase-8 was significantly associated with CEL content in the stalk via gene-based association studies. Maize lines carrying the “deletion” allele in the promoter region of Zm00001d019317 exhibited significantly higher CEL content compared to those with the “C” allele. Our study helps to understand the variation mechanism of cell wall component contents in maize stalks and contributes to genetic modification of maize resistance to stalk lodging.
format Article
id doaj-art-abd8405c578449e781eca4fb76d1f8f2
institution Kabale University
issn 1471-2164
language English
publishDate 2025-07-01
publisher BMC
record_format Article
series BMC Genomics
spelling doaj-art-abd8405c578449e781eca4fb76d1f8f22025-08-20T03:37:19ZengBMCBMC Genomics1471-21642025-07-0126111010.1186/s12864-025-11816-2Transcriptome and co-expression network analysis reveal the genetic basis of cell wall components in maize stalksYao He0Zhike Deng1Shijiang He2Zhaojie Qi3Hengshuo Chang4Peng Liu5Zhong Chen6Chaoying Zou7Yaou Shen8Langlang Ma9State Key Laboratory of Crop Gene Exploration and Utilization in Southwest China, Maize Research Institute, Sichuan Agricultural UniversityState Key Laboratory of Crop Gene Exploration and Utilization in Southwest China, Maize Research Institute, Sichuan Agricultural UniversityState Key Laboratory of Crop Gene Exploration and Utilization in Southwest China, Maize Research Institute, Sichuan Agricultural UniversityState Key Laboratory of Crop Gene Exploration and Utilization in Southwest China, Maize Research Institute, Sichuan Agricultural UniversityState Key Laboratory of Crop Gene Exploration and Utilization in Southwest China, Maize Research Institute, Sichuan Agricultural UniversityState Key Laboratory of Crop Gene Exploration and Utilization in Southwest China, Maize Research Institute, Sichuan Agricultural UniversityState Key Laboratory of Crop Gene Exploration and Utilization in Southwest China, Maize Research Institute, Sichuan Agricultural UniversityState Key Laboratory of Crop Gene Exploration and Utilization in Southwest China, Maize Research Institute, Sichuan Agricultural UniversityState Key Laboratory of Crop Gene Exploration and Utilization in Southwest China, Maize Research Institute, Sichuan Agricultural UniversityState Key Laboratory of Crop Gene Exploration and Utilization in Southwest China, Maize Research Institute, Sichuan Agricultural UniversityAbstract Cellulose (CEL), hemicellulose (HCE), and lignin (LIG) are crucial constituents of the cell wall in maize stalks, significantly influencing stalk resistance to lodging and external stresses. The proportion of these contents also determines the application of maize stalks in animal feed, energy production, and other fields. Herein, two maize inbred lines with contrasting cell wall component contents in the stalk, MO190 (high CEL, HCE, and LIG contents) and MO194 (low CEL, HCE, and LIG contents), were subjected to analysis of transcriptomes at the seedling, jointing, silking, and grain stages. By comparing the differentially expressed genes (DEGs) during stalk development between the two lines, 11,546 genes were identified as the specific DEGs in each line. Using traits-associated WGCNA (weight gene co-expression network analysis), we identified a hub co-expression module and 26 key genes regulating the CEL, HCE, and LIG contents in the stalk. Based on the eigengene-based connectivity value in the module, the four genes Zm00001d019317, Zm00001d027313, Zm00001d043965, and Zm00001d019149 were confirmed as the hub genes affecting these contents. Among these, Zm00001d019317 that encodes CEL synthase-8 was significantly associated with CEL content in the stalk via gene-based association studies. Maize lines carrying the “deletion” allele in the promoter region of Zm00001d019317 exhibited significantly higher CEL content compared to those with the “C” allele. Our study helps to understand the variation mechanism of cell wall component contents in maize stalks and contributes to genetic modification of maize resistance to stalk lodging.https://doi.org/10.1186/s12864-025-11816-2MaizeStalkCellulose (CEL)Hemicellulose (HCE)Lignin (LIG)RNA-Seq
spellingShingle Yao He
Zhike Deng
Shijiang He
Zhaojie Qi
Hengshuo Chang
Peng Liu
Zhong Chen
Chaoying Zou
Yaou Shen
Langlang Ma
Transcriptome and co-expression network analysis reveal the genetic basis of cell wall components in maize stalks
BMC Genomics
Maize
Stalk
Cellulose (CEL)
Hemicellulose (HCE)
Lignin (LIG)
RNA-Seq
title Transcriptome and co-expression network analysis reveal the genetic basis of cell wall components in maize stalks
title_full Transcriptome and co-expression network analysis reveal the genetic basis of cell wall components in maize stalks
title_fullStr Transcriptome and co-expression network analysis reveal the genetic basis of cell wall components in maize stalks
title_full_unstemmed Transcriptome and co-expression network analysis reveal the genetic basis of cell wall components in maize stalks
title_short Transcriptome and co-expression network analysis reveal the genetic basis of cell wall components in maize stalks
title_sort transcriptome and co expression network analysis reveal the genetic basis of cell wall components in maize stalks
topic Maize
Stalk
Cellulose (CEL)
Hemicellulose (HCE)
Lignin (LIG)
RNA-Seq
url https://doi.org/10.1186/s12864-025-11816-2
work_keys_str_mv AT yaohe transcriptomeandcoexpressionnetworkanalysisrevealthegeneticbasisofcellwallcomponentsinmaizestalks
AT zhikedeng transcriptomeandcoexpressionnetworkanalysisrevealthegeneticbasisofcellwallcomponentsinmaizestalks
AT shijianghe transcriptomeandcoexpressionnetworkanalysisrevealthegeneticbasisofcellwallcomponentsinmaizestalks
AT zhaojieqi transcriptomeandcoexpressionnetworkanalysisrevealthegeneticbasisofcellwallcomponentsinmaizestalks
AT hengshuochang transcriptomeandcoexpressionnetworkanalysisrevealthegeneticbasisofcellwallcomponentsinmaizestalks
AT pengliu transcriptomeandcoexpressionnetworkanalysisrevealthegeneticbasisofcellwallcomponentsinmaizestalks
AT zhongchen transcriptomeandcoexpressionnetworkanalysisrevealthegeneticbasisofcellwallcomponentsinmaizestalks
AT chaoyingzou transcriptomeandcoexpressionnetworkanalysisrevealthegeneticbasisofcellwallcomponentsinmaizestalks
AT yaoushen transcriptomeandcoexpressionnetworkanalysisrevealthegeneticbasisofcellwallcomponentsinmaizestalks
AT langlangma transcriptomeandcoexpressionnetworkanalysisrevealthegeneticbasisofcellwallcomponentsinmaizestalks