GABA and astrocytic cholesterol determine the lipid environment of GABAAR in cultured cortical neurons

Abstract The γ-aminobutyric acid (GABA) type A receptor (GABAAR), a GABA activated pentameric chloride channel, mediates fast inhibitory neurotransmission in the brain. The lipid environment is critical for GABAAR function. How lipids regulate the channel in the cell membrane is not fully understood...

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Main Authors: Zixuan Yuan, Mahmud Arif Pavel, Scott B. Hansen
Format: Article
Language:English
Published: Nature Portfolio 2025-04-01
Series:Communications Biology
Online Access:https://doi.org/10.1038/s42003-025-08026-7
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author Zixuan Yuan
Mahmud Arif Pavel
Scott B. Hansen
author_facet Zixuan Yuan
Mahmud Arif Pavel
Scott B. Hansen
author_sort Zixuan Yuan
collection DOAJ
description Abstract The γ-aminobutyric acid (GABA) type A receptor (GABAAR), a GABA activated pentameric chloride channel, mediates fast inhibitory neurotransmission in the brain. The lipid environment is critical for GABAAR function. How lipids regulate the channel in the cell membrane is not fully understood. Here we employed super resolution imaging of lipids to demonstrate that the agonist GABA induces a rapid and reversible membrane translocation of GABAAR to phosphatidylinositol 4,5-bisphosphate (PIP2) clusters in mouse primary cortical neurons. This translocation relies on nanoscopic separation of PIP2 clusters and lipid rafts (cholesterol-dependent ganglioside clusters). In a resting state, the GABAAR associates with lipid rafts and this colocalization is enhanced by uptake of astrocytic secretions. These astrocytic secretions delay desensitization and enhance maximum current. In an Alzheimer’s Disease (AD) mouse model with high brain cholesterol, GABAAR shifts into lipid rafts. Our findings suggest cholesterol is a signaling molecule and astrocytes regulates GABAARs in neurons by secreting cholesterol. The findings have implications for treating mood disorders and AD associated with altered brain lipids.
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spelling doaj-art-a404fbabf0ae4abe8092a2a2bd37eb1e2025-08-20T03:15:10ZengNature PortfolioCommunications Biology2399-36422025-04-018111310.1038/s42003-025-08026-7GABA and astrocytic cholesterol determine the lipid environment of GABAAR in cultured cortical neuronsZixuan Yuan0Mahmud Arif Pavel1Scott B. Hansen2Department of Molecular Medicine, The Scripps Research InstituteDepartment of Molecular Medicine, The Scripps Research InstituteDepartment of Molecular Medicine, The Scripps Research InstituteAbstract The γ-aminobutyric acid (GABA) type A receptor (GABAAR), a GABA activated pentameric chloride channel, mediates fast inhibitory neurotransmission in the brain. The lipid environment is critical for GABAAR function. How lipids regulate the channel in the cell membrane is not fully understood. Here we employed super resolution imaging of lipids to demonstrate that the agonist GABA induces a rapid and reversible membrane translocation of GABAAR to phosphatidylinositol 4,5-bisphosphate (PIP2) clusters in mouse primary cortical neurons. This translocation relies on nanoscopic separation of PIP2 clusters and lipid rafts (cholesterol-dependent ganglioside clusters). In a resting state, the GABAAR associates with lipid rafts and this colocalization is enhanced by uptake of astrocytic secretions. These astrocytic secretions delay desensitization and enhance maximum current. In an Alzheimer’s Disease (AD) mouse model with high brain cholesterol, GABAAR shifts into lipid rafts. Our findings suggest cholesterol is a signaling molecule and astrocytes regulates GABAARs in neurons by secreting cholesterol. The findings have implications for treating mood disorders and AD associated with altered brain lipids.https://doi.org/10.1038/s42003-025-08026-7
spellingShingle Zixuan Yuan
Mahmud Arif Pavel
Scott B. Hansen
GABA and astrocytic cholesterol determine the lipid environment of GABAAR in cultured cortical neurons
Communications Biology
title GABA and astrocytic cholesterol determine the lipid environment of GABAAR in cultured cortical neurons
title_full GABA and astrocytic cholesterol determine the lipid environment of GABAAR in cultured cortical neurons
title_fullStr GABA and astrocytic cholesterol determine the lipid environment of GABAAR in cultured cortical neurons
title_full_unstemmed GABA and astrocytic cholesterol determine the lipid environment of GABAAR in cultured cortical neurons
title_short GABA and astrocytic cholesterol determine the lipid environment of GABAAR in cultured cortical neurons
title_sort gaba and astrocytic cholesterol determine the lipid environment of gabaar in cultured cortical neurons
url https://doi.org/10.1038/s42003-025-08026-7
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AT scottbhansen gabaandastrocyticcholesteroldeterminethelipidenvironmentofgabaarinculturedcorticalneurons