Matrix regulation: a plug-and-tune method for combinatorial regulation in Saccharomyces cerevisiae

Abstract Transcriptional fine-tuning of long pathways is complex, even in the extensively applied cell factory Saccharomyces cerevisiae. Here, we present Matrix Regulation (MR), a CRISPR-mediated pathway fine-tuning method enabling the construction of 68 gRNA combinations and screening for the optim...

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Main Authors: Xiaolong Teng, Zibai Wang, Yueping Zhang, Binhao Wang, Guiping Gong, Jinmiao Hu, Yifan Zhu, Baoyi Peng, Junyang Wang, James Chen, Shuobo Shi, Jens Nielsen, Zihe Liu
Format: Article
Language:English
Published: Nature Portfolio 2025-08-01
Series:Nature Communications
Online Access:https://doi.org/10.1038/s41467-025-62886-5
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author Xiaolong Teng
Zibai Wang
Yueping Zhang
Binhao Wang
Guiping Gong
Jinmiao Hu
Yifan Zhu
Baoyi Peng
Junyang Wang
James Chen
Shuobo Shi
Jens Nielsen
Zihe Liu
author_facet Xiaolong Teng
Zibai Wang
Yueping Zhang
Binhao Wang
Guiping Gong
Jinmiao Hu
Yifan Zhu
Baoyi Peng
Junyang Wang
James Chen
Shuobo Shi
Jens Nielsen
Zihe Liu
author_sort Xiaolong Teng
collection DOAJ
description Abstract Transcriptional fine-tuning of long pathways is complex, even in the extensively applied cell factory Saccharomyces cerevisiae. Here, we present Matrix Regulation (MR), a CRISPR-mediated pathway fine-tuning method enabling the construction of 68 gRNA combinations and screening for the optimal expression levels across up to eight genes. We first identify multiple tRNAs with efficient gRNA processing capacities to assemble a gRNA regulatory matrix combinatorially. Then, we expand the target recognition of CRISPR regulation from NGG PAM to NG PAM by characterizing dCas9 variants. To increase the dynamic range of modulation, we test 101 candidate activation domains followed by mutagenesis and screening the best one to further enhance its activation capability in S. cerevisiae by 3-fold. The regulations generate combinatorial strain libraries for both the mevalonate pathway and the heme biosynthesis pathway and increase squalene production by 37-fold and heme by 17-fold, respectively, demonstrating the versatility of our method and its applicability in fundamental research.
format Article
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institution Kabale University
issn 2041-1723
language English
publishDate 2025-08-01
publisher Nature Portfolio
record_format Article
series Nature Communications
spelling doaj-art-a1ea37403ee74c7886914b4b765328df2025-08-20T03:43:16ZengNature PortfolioNature Communications2041-17232025-08-0116111210.1038/s41467-025-62886-5Matrix regulation: a plug-and-tune method for combinatorial regulation in Saccharomyces cerevisiaeXiaolong Teng0Zibai Wang1Yueping Zhang2Binhao Wang3Guiping Gong4Jinmiao Hu5Yifan Zhu6Baoyi Peng7Junyang Wang8James Chen9Shuobo Shi10Jens Nielsen11Zihe Liu12College of Life Science and Technology, State Key Laboratory of Green Biomanufacturing, Beijing Advanced Innovation Center for Soft Matter Science and Engineering, Beijing University of Chemical TechnologyCollege of Life Science and Technology, State Key Laboratory of Green Biomanufacturing, Beijing Advanced Innovation Center for Soft Matter Science and Engineering, Beijing University of Chemical TechnologyCollege of Veterinary Medicine, China Agricultural UniversityCollege of Life Science and Technology, State Key Laboratory of Green Biomanufacturing, Beijing Advanced Innovation Center for Soft Matter Science and Engineering, Beijing University of Chemical TechnologyCollege of Life Science and Technology, State Key Laboratory of Green Biomanufacturing, Beijing Advanced Innovation Center for Soft Matter Science and Engineering, Beijing University of Chemical TechnologyCollege of Life Science and Technology, State Key Laboratory of Green Biomanufacturing, Beijing Advanced Innovation Center for Soft Matter Science and Engineering, Beijing University of Chemical TechnologyCollege of Life Science and Technology, State Key Laboratory of Green Biomanufacturing, Beijing Advanced Innovation Center for Soft Matter Science and Engineering, Beijing University of Chemical TechnologyCollege of Life Science and Technology, State Key Laboratory of Green Biomanufacturing, Beijing Advanced Innovation Center for Soft Matter Science and Engineering, Beijing University of Chemical TechnologyCollege of Life Science and Technology, State Key Laboratory of Green Biomanufacturing, Beijing Advanced Innovation Center for Soft Matter Science and Engineering, Beijing University of Chemical TechnologyCollege of Life Science and Technology, State Key Laboratory of Green Biomanufacturing, Beijing Advanced Innovation Center for Soft Matter Science and Engineering, Beijing University of Chemical TechnologyCollege of Life Science and Technology, State Key Laboratory of Green Biomanufacturing, Beijing Advanced Innovation Center for Soft Matter Science and Engineering, Beijing University of Chemical TechnologyCollege of Life Science and Technology, State Key Laboratory of Green Biomanufacturing, Beijing Advanced Innovation Center for Soft Matter Science and Engineering, Beijing University of Chemical TechnologyCollege of Life Science and Technology, State Key Laboratory of Green Biomanufacturing, Beijing Advanced Innovation Center for Soft Matter Science and Engineering, Beijing University of Chemical TechnologyAbstract Transcriptional fine-tuning of long pathways is complex, even in the extensively applied cell factory Saccharomyces cerevisiae. Here, we present Matrix Regulation (MR), a CRISPR-mediated pathway fine-tuning method enabling the construction of 68 gRNA combinations and screening for the optimal expression levels across up to eight genes. We first identify multiple tRNAs with efficient gRNA processing capacities to assemble a gRNA regulatory matrix combinatorially. Then, we expand the target recognition of CRISPR regulation from NGG PAM to NG PAM by characterizing dCas9 variants. To increase the dynamic range of modulation, we test 101 candidate activation domains followed by mutagenesis and screening the best one to further enhance its activation capability in S. cerevisiae by 3-fold. The regulations generate combinatorial strain libraries for both the mevalonate pathway and the heme biosynthesis pathway and increase squalene production by 37-fold and heme by 17-fold, respectively, demonstrating the versatility of our method and its applicability in fundamental research.https://doi.org/10.1038/s41467-025-62886-5
spellingShingle Xiaolong Teng
Zibai Wang
Yueping Zhang
Binhao Wang
Guiping Gong
Jinmiao Hu
Yifan Zhu
Baoyi Peng
Junyang Wang
James Chen
Shuobo Shi
Jens Nielsen
Zihe Liu
Matrix regulation: a plug-and-tune method for combinatorial regulation in Saccharomyces cerevisiae
Nature Communications
title Matrix regulation: a plug-and-tune method for combinatorial regulation in Saccharomyces cerevisiae
title_full Matrix regulation: a plug-and-tune method for combinatorial regulation in Saccharomyces cerevisiae
title_fullStr Matrix regulation: a plug-and-tune method for combinatorial regulation in Saccharomyces cerevisiae
title_full_unstemmed Matrix regulation: a plug-and-tune method for combinatorial regulation in Saccharomyces cerevisiae
title_short Matrix regulation: a plug-and-tune method for combinatorial regulation in Saccharomyces cerevisiae
title_sort matrix regulation a plug and tune method for combinatorial regulation in saccharomyces cerevisiae
url https://doi.org/10.1038/s41467-025-62886-5
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