Programming virulent bacteriophages by developing a multiplex genome engineering method

ABSTRACT The use of virulent bacteriophages (phages) against pathogenic bacteria has recently attracted considerable interest. The limitations of naturally isolated phages have promoted the development of genome engineering methods to optimize their functions; however, engineering of virulent phage...

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Main Authors: Hailin Zhang, Ru Zhu, Zhaofei Wang, Ruoting He, Yuran Zhang, Ji Luan, Yaxian Yan, Youming Zhang, Hailong Wang
Format: Article
Language:English
Published: American Society for Microbiology 2025-06-01
Series:mBio
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Online Access:https://journals.asm.org/doi/10.1128/mbio.03582-24
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author Hailin Zhang
Ru Zhu
Zhaofei Wang
Ruoting He
Yuran Zhang
Ji Luan
Yaxian Yan
Youming Zhang
Hailong Wang
author_facet Hailin Zhang
Ru Zhu
Zhaofei Wang
Ruoting He
Yuran Zhang
Ji Luan
Yaxian Yan
Youming Zhang
Hailong Wang
author_sort Hailin Zhang
collection DOAJ
description ABSTRACT The use of virulent bacteriophages (phages) against pathogenic bacteria has recently attracted considerable interest. The limitations of naturally isolated phages have promoted the development of genome engineering methods to optimize their functions; however, engineering of virulent phage genomes in bacterial hosts remains challenging. Here, we describe a SMART (splitting, modifying, assembling, and rebooting) method for multiplex genome engineering of virulent phages by splitting the genome into multiple segments cloned and inserted into single-copy bacterial artificial chromosome vectors in Escherichia coli to overcome the toxicity of phage gene products, modifying multiple targeted loci in parallel through recombineering, assembling split segments into an intact genome, and rebooting the recombined phage in bacterial hosts. Using the SMART method, the 39.9-kb T7 E. coli phage genome was split into 10 segments, and 8 genomic loci were deleted in parallel to obtain a chassis phage with 10% genome reduction. The insertion capacity of the chassis phage genome and the expression levels of exogenous genes at different loci were evaluated. Finally, a synthetic T7 phage that efficiently lyses widely different bacteria, such as E. coli, Staphylococcus aureus, and Streptococcus agalactiae, was constructed by expressing heterologous lysins from Staphylococcus and Streptococcus phages. SMART will facilitate the programming and understanding of the genome structure and function of virulent phages.IMPORTANCEUnlike temperate phages, which integrate into host genomes and allow time for genetic manipulation, lytic phages rapidly hijack the bacterial machinery and trigger host lysis within minutes, leaving an extremely narrow editing window. Furthermore, virulent phage genes usually encode toxic products that inhibit the growth of bacterial hosts. We developed a SMART (splitting, modifying, assembling, and rebooting) method for multiplex genome engineering of virulent phages. We deleted 3.9 kb sequences distributed across 8 sites in the 39.9 kb genome of the wild-type T7 Escherichia coli phage to construct a chassis phage. Synthetic T7 phages expressing heterologous lysin genes were constructed to efficiently lyse different bacteria, such as E. coli, Staphylococcus aureus, or Streptococcus agalactiae. The SMART method will facilitate targeted modifications of phage genomes for the creation of custom-designed phages with enhanced therapeutic efficacy, broader host specificity, and programmable behaviors.
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spelling doaj-art-a16f958d0e854b28b2ac09fd429df7ef2025-08-20T03:10:50ZengAmerican Society for MicrobiologymBio2150-75112025-06-0116610.1128/mbio.03582-24Programming virulent bacteriophages by developing a multiplex genome engineering methodHailin Zhang0Ru Zhu1Zhaofei Wang2Ruoting He3Yuran Zhang4Ji Luan5Yaxian Yan6Youming Zhang7Hailong Wang8State Key Laboratory of Microbial Technology, Institute of Microbial Technology, Helmholtz International Lab for Anti-infectives, Shandong University–Helmholtz Institute of Biotechnology, Shandong University, Qingdao, Shandong, ChinaState Key Laboratory of Microbial Technology, Institute of Microbial Technology, Helmholtz International Lab for Anti-infectives, Shandong University–Helmholtz Institute of Biotechnology, Shandong University, Qingdao, Shandong, ChinaSchool of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai Key Laboratory of Veterinary Biotechnology, Shanghai, ChinaState Key Laboratory of Microbial Technology, Institute of Microbial Technology, Helmholtz International Lab for Anti-infectives, Shandong University–Helmholtz Institute of Biotechnology, Shandong University, Qingdao, Shandong, ChinaSchool of Life Sciences, Jining Medical University, Rizhao, Shandong, ChinaState Key Laboratory of Microbial Technology, Institute of Microbial Technology, Helmholtz International Lab for Anti-infectives, Shandong University–Helmholtz Institute of Biotechnology, Shandong University, Qingdao, Shandong, ChinaSchool of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai Key Laboratory of Veterinary Biotechnology, Shanghai, ChinaState Key Laboratory of Microbial Technology, Institute of Microbial Technology, Helmholtz International Lab for Anti-infectives, Shandong University–Helmholtz Institute of Biotechnology, Shandong University, Qingdao, Shandong, ChinaState Key Laboratory of Microbial Technology, Institute of Microbial Technology, Helmholtz International Lab for Anti-infectives, Shandong University–Helmholtz Institute of Biotechnology, Shandong University, Qingdao, Shandong, ChinaABSTRACT The use of virulent bacteriophages (phages) against pathogenic bacteria has recently attracted considerable interest. The limitations of naturally isolated phages have promoted the development of genome engineering methods to optimize their functions; however, engineering of virulent phage genomes in bacterial hosts remains challenging. Here, we describe a SMART (splitting, modifying, assembling, and rebooting) method for multiplex genome engineering of virulent phages by splitting the genome into multiple segments cloned and inserted into single-copy bacterial artificial chromosome vectors in Escherichia coli to overcome the toxicity of phage gene products, modifying multiple targeted loci in parallel through recombineering, assembling split segments into an intact genome, and rebooting the recombined phage in bacterial hosts. Using the SMART method, the 39.9-kb T7 E. coli phage genome was split into 10 segments, and 8 genomic loci were deleted in parallel to obtain a chassis phage with 10% genome reduction. The insertion capacity of the chassis phage genome and the expression levels of exogenous genes at different loci were evaluated. Finally, a synthetic T7 phage that efficiently lyses widely different bacteria, such as E. coli, Staphylococcus aureus, and Streptococcus agalactiae, was constructed by expressing heterologous lysins from Staphylococcus and Streptococcus phages. SMART will facilitate the programming and understanding of the genome structure and function of virulent phages.IMPORTANCEUnlike temperate phages, which integrate into host genomes and allow time for genetic manipulation, lytic phages rapidly hijack the bacterial machinery and trigger host lysis within minutes, leaving an extremely narrow editing window. Furthermore, virulent phage genes usually encode toxic products that inhibit the growth of bacterial hosts. We developed a SMART (splitting, modifying, assembling, and rebooting) method for multiplex genome engineering of virulent phages. We deleted 3.9 kb sequences distributed across 8 sites in the 39.9 kb genome of the wild-type T7 Escherichia coli phage to construct a chassis phage. Synthetic T7 phages expressing heterologous lysin genes were constructed to efficiently lyse different bacteria, such as E. coli, Staphylococcus aureus, or Streptococcus agalactiae. The SMART method will facilitate targeted modifications of phage genomes for the creation of custom-designed phages with enhanced therapeutic efficacy, broader host specificity, and programmable behaviors.https://journals.asm.org/doi/10.1128/mbio.03582-24phage engineeringsynthetic phagesgenome engineeringrecombineeringphage chassis
spellingShingle Hailin Zhang
Ru Zhu
Zhaofei Wang
Ruoting He
Yuran Zhang
Ji Luan
Yaxian Yan
Youming Zhang
Hailong Wang
Programming virulent bacteriophages by developing a multiplex genome engineering method
mBio
phage engineering
synthetic phages
genome engineering
recombineering
phage chassis
title Programming virulent bacteriophages by developing a multiplex genome engineering method
title_full Programming virulent bacteriophages by developing a multiplex genome engineering method
title_fullStr Programming virulent bacteriophages by developing a multiplex genome engineering method
title_full_unstemmed Programming virulent bacteriophages by developing a multiplex genome engineering method
title_short Programming virulent bacteriophages by developing a multiplex genome engineering method
title_sort programming virulent bacteriophages by developing a multiplex genome engineering method
topic phage engineering
synthetic phages
genome engineering
recombineering
phage chassis
url https://journals.asm.org/doi/10.1128/mbio.03582-24
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