Host-Specific and Environment-Dependent Effects of Endophyte <i>Alternaria oxytropis</i> on Three Locoweed <i>Oxytropis</i> Species in China

Plant–endophyte symbioses are widespread in grasslands. While symbiotic interactions often provide hosts with major fitness enhancements, the role of the endophyte <i>Alternaria oxytropis</i>, which produces swainsonine in locoweeds (<i>Oxytropis</i> and <i>Astragalus&l...

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Main Authors: Yue-Yang Zhang, Yan-Zhong Li, Zun-Ji Shi
Format: Article
Language:English
Published: MDPI AG 2025-07-01
Series:Journal of Fungi
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Online Access:https://www.mdpi.com/2309-608X/11/7/516
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author Yue-Yang Zhang
Yan-Zhong Li
Zun-Ji Shi
author_facet Yue-Yang Zhang
Yan-Zhong Li
Zun-Ji Shi
author_sort Yue-Yang Zhang
collection DOAJ
description Plant–endophyte symbioses are widespread in grasslands. While symbiotic interactions often provide hosts with major fitness enhancements, the role of the endophyte <i>Alternaria oxytropis</i>, which produces swainsonine in locoweeds (<i>Oxytropis</i> and <i>Astragalus</i> spp.), remains enigmatic. We compared endophyte-infected (E+) and endophyte-free (E−) plants of three main Chinese locoweed species (<i>O. kansuensis</i>, <i>O. glabra</i>, and <i>O. ochrocephala</i>) under controlled conditions, and analyzed environmental factors at locoweed poisoning hotspots for herbivores. The results demonstrated significant species-specific effects: E+ plants of <i>O. glabra</i> and <i>O. ochrocephala</i> exhibited 26–39% reductions in biomass, net photosynthetic rate, and stomatal conductance, with elevated CO<sub>2</sub> levels, while <i>O. kansuensis</i> showed no measurable impacts. Swainsonine concentrations were 16–20 times higher in E+ plants (122.6–151.7 mg/kg) than in E− plants. Geospatial analysis revealed that poisoning hotspots for herbivores consistently occurred in regions with extreme winter conditions (minimum temperatures ≤ −17 °C and precipitation ≤ 1 mm during the driest month), suggesting context-dependent benefits under abiotic stress. These findings suggest that the ecological role of <i>A. oxytropis</i> may vary depending on both host species and environmental context, highlighting a trade-off between growth costs and potential stress tolerance conferred by <i>A. oxytropis</i>. The study underscores the need for field validation to elucidate the adaptive mechanisms maintaining this symbiosis in harsh environments.
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spelling doaj-art-9fe32ece64d642eda6cdb2e7731a92ae2025-08-20T03:08:09ZengMDPI AGJournal of Fungi2309-608X2025-07-0111751610.3390/jof11070516Host-Specific and Environment-Dependent Effects of Endophyte <i>Alternaria oxytropis</i> on Three Locoweed <i>Oxytropis</i> Species in ChinaYue-Yang Zhang0Yan-Zhong Li1Zun-Ji Shi2State Key Laboratory of Herbage Improvement and Grassland Agro-Ecosystems, College of Pastoral Agriculture Science and Technology, Center for Grassland Microbiome, Engineering Research Center of Grassland Industry, Ministry of Education, Lanzhou University, Lanzhou 730020, ChinaState Key Laboratory of Herbage Improvement and Grassland Agro-Ecosystems, College of Pastoral Agriculture Science and Technology, Center for Grassland Microbiome, Engineering Research Center of Grassland Industry, Ministry of Education, Lanzhou University, Lanzhou 730020, ChinaState Key Laboratory of Herbage Improvement and Grassland Agro-Ecosystems, College of Pastoral Agriculture Science and Technology, Center for Grassland Microbiome, Engineering Research Center of Grassland Industry, Ministry of Education, Lanzhou University, Lanzhou 730020, ChinaPlant–endophyte symbioses are widespread in grasslands. While symbiotic interactions often provide hosts with major fitness enhancements, the role of the endophyte <i>Alternaria oxytropis</i>, which produces swainsonine in locoweeds (<i>Oxytropis</i> and <i>Astragalus</i> spp.), remains enigmatic. We compared endophyte-infected (E+) and endophyte-free (E−) plants of three main Chinese locoweed species (<i>O. kansuensis</i>, <i>O. glabra</i>, and <i>O. ochrocephala</i>) under controlled conditions, and analyzed environmental factors at locoweed poisoning hotspots for herbivores. The results demonstrated significant species-specific effects: E+ plants of <i>O. glabra</i> and <i>O. ochrocephala</i> exhibited 26–39% reductions in biomass, net photosynthetic rate, and stomatal conductance, with elevated CO<sub>2</sub> levels, while <i>O. kansuensis</i> showed no measurable impacts. Swainsonine concentrations were 16–20 times higher in E+ plants (122.6–151.7 mg/kg) than in E− plants. Geospatial analysis revealed that poisoning hotspots for herbivores consistently occurred in regions with extreme winter conditions (minimum temperatures ≤ −17 °C and precipitation ≤ 1 mm during the driest month), suggesting context-dependent benefits under abiotic stress. These findings suggest that the ecological role of <i>A. oxytropis</i> may vary depending on both host species and environmental context, highlighting a trade-off between growth costs and potential stress tolerance conferred by <i>A. oxytropis</i>. The study underscores the need for field validation to elucidate the adaptive mechanisms maintaining this symbiosis in harsh environments.https://www.mdpi.com/2309-608X/11/7/516endosymbiontphotosynthesisswainsoninelocoweedabiotic stress
spellingShingle Yue-Yang Zhang
Yan-Zhong Li
Zun-Ji Shi
Host-Specific and Environment-Dependent Effects of Endophyte <i>Alternaria oxytropis</i> on Three Locoweed <i>Oxytropis</i> Species in China
Journal of Fungi
endosymbiont
photosynthesis
swainsonine
locoweed
abiotic stress
title Host-Specific and Environment-Dependent Effects of Endophyte <i>Alternaria oxytropis</i> on Three Locoweed <i>Oxytropis</i> Species in China
title_full Host-Specific and Environment-Dependent Effects of Endophyte <i>Alternaria oxytropis</i> on Three Locoweed <i>Oxytropis</i> Species in China
title_fullStr Host-Specific and Environment-Dependent Effects of Endophyte <i>Alternaria oxytropis</i> on Three Locoweed <i>Oxytropis</i> Species in China
title_full_unstemmed Host-Specific and Environment-Dependent Effects of Endophyte <i>Alternaria oxytropis</i> on Three Locoweed <i>Oxytropis</i> Species in China
title_short Host-Specific and Environment-Dependent Effects of Endophyte <i>Alternaria oxytropis</i> on Three Locoweed <i>Oxytropis</i> Species in China
title_sort host specific and environment dependent effects of endophyte i alternaria oxytropis i on three locoweed i oxytropis i species in china
topic endosymbiont
photosynthesis
swainsonine
locoweed
abiotic stress
url https://www.mdpi.com/2309-608X/11/7/516
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AT yanzhongli hostspecificandenvironmentdependenteffectsofendophyteialternariaoxytropisionthreelocoweedioxytropisispeciesinchina
AT zunjishi hostspecificandenvironmentdependenteffectsofendophyteialternariaoxytropisionthreelocoweedioxytropisispeciesinchina