Influence of nitrate-containing arugula juice on nitrate-reducing oral bacteria and periodontopathogens in smokers’ biofilm

Green leafy vegetables such as arugula are rich in nitrates that support oral health, yet their effects on oral microbial balance, especially in smokers, remain unclear. This study evaluated the survival and activity of nitrate-reducing bacteria (NRB; Veillonella spp. and Rothia spp.) in biofilm exp...

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Main Authors: Boy M. Bachtiar, Irene E. Rieuwpassa, Heni Susilowati, Retno Indrawati, Citra F. Theodorea, Turmidzi Fath, Endang W. Bachtiar
Format: Article
Language:English
Published: Frontiers Media S.A. 2025-05-01
Series:Frontiers in Dental Medicine
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Online Access:https://www.frontiersin.org/articles/10.3389/fdmed.2025.1545479/full
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author Boy M. Bachtiar
Boy M. Bachtiar
Irene E. Rieuwpassa
Heni Susilowati
Retno Indrawati
Citra F. Theodorea
Citra F. Theodorea
Turmidzi Fath
Endang W. Bachtiar
Endang W. Bachtiar
author_facet Boy M. Bachtiar
Boy M. Bachtiar
Irene E. Rieuwpassa
Heni Susilowati
Retno Indrawati
Citra F. Theodorea
Citra F. Theodorea
Turmidzi Fath
Endang W. Bachtiar
Endang W. Bachtiar
author_sort Boy M. Bachtiar
collection DOAJ
description Green leafy vegetables such as arugula are rich in nitrates that support oral health, yet their effects on oral microbial balance, especially in smokers, remain unclear. This study evaluated the survival and activity of nitrate-reducing bacteria (NRB; Veillonella spp. and Rothia spp.) in biofilm exposed to nitrate-containing arugula juice (3.25 or 6.25 μM). The proportions of NRB were compared with periodontopathogens (Porphyromonas gingivalis and Fusobacterium nucleatum). Using quantitative real-time PCR (qPCR), we assessed bacterial survival and transcription of nitrate reductase genes (narG and napA) in biofilm from smokers and non-smokers. The results revealed that nitrate-containing arugula juice increased NRB bacteria abundance while reducing periodontopathogen growth. A higher level of nitrate (6.25 μM) increased nitrate reductase expression. Prolonged exposure (9 h) sustained the growth-promoting effect on Rothia spp. These results suggest that non-smokers have more nitrate-reducing bacteria in their biofilm, which promotes oral microbial balance. Thus, smokers might be advised to consume nitrate-containing arugula juice to promote NRB, which may have health benefits.
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series Frontiers in Dental Medicine
spelling doaj-art-9c08db792dd14828b6f41a6783dba8852025-08-20T02:16:12ZengFrontiers Media S.A.Frontiers in Dental Medicine2673-49152025-05-01610.3389/fdmed.2025.15454791545479Influence of nitrate-containing arugula juice on nitrate-reducing oral bacteria and periodontopathogens in smokers’ biofilmBoy M. Bachtiar0Boy M. Bachtiar1Irene E. Rieuwpassa2Heni Susilowati3Retno Indrawati4Citra F. Theodorea5Citra F. Theodorea6Turmidzi Fath7Endang W. Bachtiar8Endang W. Bachtiar9Department of Oral Biology, Faculty of Dentistry, Universitas Indonesia, Jakarta, IndonesiaOral Science Research Center, Faculty of Dentistry, Universitas Indonesia, Jakarta, IndonesiaDepartment of Oral Biology, Faculty of Dentistry, Universitas Hasanuddin, Makassar, IndonesiaDepartment of Oral Biology, Faculty of Dentistry, Universitas Gadjah Mada, Yokyakarta, IndonesiaDepartment of Oral Biology, Dental Medicine Faculty, Universitas Airlangga, Surabaya, IndonesiaDepartment of Oral Biology, Faculty of Dentistry, Universitas Indonesia, Jakarta, IndonesiaOral Science Research Center, Faculty of Dentistry, Universitas Indonesia, Jakarta, IndonesiaOral Science Research Center, Faculty of Dentistry, Universitas Indonesia, Jakarta, IndonesiaDepartment of Oral Biology, Faculty of Dentistry, Universitas Indonesia, Jakarta, IndonesiaOral Science Research Center, Faculty of Dentistry, Universitas Indonesia, Jakarta, IndonesiaGreen leafy vegetables such as arugula are rich in nitrates that support oral health, yet their effects on oral microbial balance, especially in smokers, remain unclear. This study evaluated the survival and activity of nitrate-reducing bacteria (NRB; Veillonella spp. and Rothia spp.) in biofilm exposed to nitrate-containing arugula juice (3.25 or 6.25 μM). The proportions of NRB were compared with periodontopathogens (Porphyromonas gingivalis and Fusobacterium nucleatum). Using quantitative real-time PCR (qPCR), we assessed bacterial survival and transcription of nitrate reductase genes (narG and napA) in biofilm from smokers and non-smokers. The results revealed that nitrate-containing arugula juice increased NRB bacteria abundance while reducing periodontopathogen growth. A higher level of nitrate (6.25 μM) increased nitrate reductase expression. Prolonged exposure (9 h) sustained the growth-promoting effect on Rothia spp. These results suggest that non-smokers have more nitrate-reducing bacteria in their biofilm, which promotes oral microbial balance. Thus, smokers might be advised to consume nitrate-containing arugula juice to promote NRB, which may have health benefits.https://www.frontiersin.org/articles/10.3389/fdmed.2025.1545479/fullarugulabiofilmnitrate-reducing bacteriaperiodontopathogensnitrate-associated genes
spellingShingle Boy M. Bachtiar
Boy M. Bachtiar
Irene E. Rieuwpassa
Heni Susilowati
Retno Indrawati
Citra F. Theodorea
Citra F. Theodorea
Turmidzi Fath
Endang W. Bachtiar
Endang W. Bachtiar
Influence of nitrate-containing arugula juice on nitrate-reducing oral bacteria and periodontopathogens in smokers’ biofilm
Frontiers in Dental Medicine
arugula
biofilm
nitrate-reducing bacteria
periodontopathogens
nitrate-associated genes
title Influence of nitrate-containing arugula juice on nitrate-reducing oral bacteria and periodontopathogens in smokers’ biofilm
title_full Influence of nitrate-containing arugula juice on nitrate-reducing oral bacteria and periodontopathogens in smokers’ biofilm
title_fullStr Influence of nitrate-containing arugula juice on nitrate-reducing oral bacteria and periodontopathogens in smokers’ biofilm
title_full_unstemmed Influence of nitrate-containing arugula juice on nitrate-reducing oral bacteria and periodontopathogens in smokers’ biofilm
title_short Influence of nitrate-containing arugula juice on nitrate-reducing oral bacteria and periodontopathogens in smokers’ biofilm
title_sort influence of nitrate containing arugula juice on nitrate reducing oral bacteria and periodontopathogens in smokers biofilm
topic arugula
biofilm
nitrate-reducing bacteria
periodontopathogens
nitrate-associated genes
url https://www.frontiersin.org/articles/10.3389/fdmed.2025.1545479/full
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