Transcriptome analysis and genome-wide identification of WRKY gene family in Leonurus japonicus under drought stress

Abstract Background Leonurus japonicus (L. japonicus) is a herbaceous flowering plant, widely distributed in Asia. Drought is one of the primary environmental stress factors affecting L. japonicus growth. Previous studies have demonstrated that WRKY transcription factors (TFs) play a crucial role in...

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Main Authors: Tongtong Guo, Qing Wang, Yuxiao Yi, Zelin Yin, Lili Xu, Fengtang Yang, Jianing Xu, Ming Zhang
Format: Article
Language:English
Published: BMC 2025-05-01
Series:BMC Plant Biology
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Online Access:https://doi.org/10.1186/s12870-025-06606-7
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author Tongtong Guo
Qing Wang
Yuxiao Yi
Zelin Yin
Lili Xu
Fengtang Yang
Jianing Xu
Ming Zhang
author_facet Tongtong Guo
Qing Wang
Yuxiao Yi
Zelin Yin
Lili Xu
Fengtang Yang
Jianing Xu
Ming Zhang
author_sort Tongtong Guo
collection DOAJ
description Abstract Background Leonurus japonicus (L. japonicus) is a herbaceous flowering plant, widely distributed in Asia. Drought is one of the primary environmental stress factors affecting L. japonicus growth. Previous studies have demonstrated that WRKY transcription factors (TFs) play a crucial role in plant responses to drought stress. So far, there has been no research on the function of WRKY genes in L. japonicus. Results The physiological experiment results showed that drought stress significantly increased the malondialdehyde (MDA), proline, and hydrogen peroxide (H₂O₂) content of L. japonicus. Transcriptome analysis revealed significant changes in the expression levels of the WRKY gene family. Based on bioinformatics analysis, 67 WRKY genes (LjWRKYs) were identified in the genome of L. japonicus, with amino acid lengths ranging from 85 to 574. The LjWRKYs can be divided into three subfamilies. Among them, the expression of LjWRKY (1/4/23/44) were significantly up-regulated under drought stress, whereas the expression of LjWRKY (21/25/65) were significantly down-regulated. Additionally, Kyoto Encyclopedia of Genes and Genomes (KEGG) analysis indicated that after drought stress, differentially expressed genes (DEGs) were enriched in plant hormone signal transduction pathway, the MAPK signaling pathway and biosynthesis of secondary metabolites pathway. In the MAPK pathway, there were 19 DEGs, 9 of which contained W-box regions, suggesting that they may be potential regulatory targets of LjWRKY TFs under drought stress. Conclusion These findings suggested that WRKY gene family may participate in the response to drought stress in L. japonicus. This study provides a scientific basis for the further development and functional validation of the WRKY gene family in L. japonicus. Clinical trial number Not applicable.
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spelling doaj-art-9b491a24ae4845c387ab3280a79772da2025-08-20T01:49:36ZengBMCBMC Plant Biology1471-22292025-05-0125111910.1186/s12870-025-06606-7Transcriptome analysis and genome-wide identification of WRKY gene family in Leonurus japonicus under drought stressTongtong Guo0Qing Wang1Yuxiao Yi2Zelin Yin3Lili Xu4Fengtang Yang5Jianing Xu6Ming Zhang7School of Life Sciences and Medicine, Shandong University of TechnologySchool of Life Sciences and Medicine, Shandong University of TechnologySchool of Life Sciences and Medicine, Shandong University of TechnologySchool of Life Sciences and Medicine, Shandong University of TechnologySchool of Life Sciences and Medicine, Shandong University of TechnologySchool of Life Sciences and Medicine, Shandong University of TechnologySchool of Life Sciences and Medicine, Shandong University of TechnologyCollege of Agriculture and Bioengineering (Peony College), Heze UniversityAbstract Background Leonurus japonicus (L. japonicus) is a herbaceous flowering plant, widely distributed in Asia. Drought is one of the primary environmental stress factors affecting L. japonicus growth. Previous studies have demonstrated that WRKY transcription factors (TFs) play a crucial role in plant responses to drought stress. So far, there has been no research on the function of WRKY genes in L. japonicus. Results The physiological experiment results showed that drought stress significantly increased the malondialdehyde (MDA), proline, and hydrogen peroxide (H₂O₂) content of L. japonicus. Transcriptome analysis revealed significant changes in the expression levels of the WRKY gene family. Based on bioinformatics analysis, 67 WRKY genes (LjWRKYs) were identified in the genome of L. japonicus, with amino acid lengths ranging from 85 to 574. The LjWRKYs can be divided into three subfamilies. Among them, the expression of LjWRKY (1/4/23/44) were significantly up-regulated under drought stress, whereas the expression of LjWRKY (21/25/65) were significantly down-regulated. Additionally, Kyoto Encyclopedia of Genes and Genomes (KEGG) analysis indicated that after drought stress, differentially expressed genes (DEGs) were enriched in plant hormone signal transduction pathway, the MAPK signaling pathway and biosynthesis of secondary metabolites pathway. In the MAPK pathway, there were 19 DEGs, 9 of which contained W-box regions, suggesting that they may be potential regulatory targets of LjWRKY TFs under drought stress. Conclusion These findings suggested that WRKY gene family may participate in the response to drought stress in L. japonicus. This study provides a scientific basis for the further development and functional validation of the WRKY gene family in L. japonicus. Clinical trial number Not applicable.https://doi.org/10.1186/s12870-025-06606-7Leonurus japonicusWRKY familyGenome-wide analysisDrought stress
spellingShingle Tongtong Guo
Qing Wang
Yuxiao Yi
Zelin Yin
Lili Xu
Fengtang Yang
Jianing Xu
Ming Zhang
Transcriptome analysis and genome-wide identification of WRKY gene family in Leonurus japonicus under drought stress
BMC Plant Biology
Leonurus japonicus
WRKY family
Genome-wide analysis
Drought stress
title Transcriptome analysis and genome-wide identification of WRKY gene family in Leonurus japonicus under drought stress
title_full Transcriptome analysis and genome-wide identification of WRKY gene family in Leonurus japonicus under drought stress
title_fullStr Transcriptome analysis and genome-wide identification of WRKY gene family in Leonurus japonicus under drought stress
title_full_unstemmed Transcriptome analysis and genome-wide identification of WRKY gene family in Leonurus japonicus under drought stress
title_short Transcriptome analysis and genome-wide identification of WRKY gene family in Leonurus japonicus under drought stress
title_sort transcriptome analysis and genome wide identification of wrky gene family in leonurus japonicus under drought stress
topic Leonurus japonicus
WRKY family
Genome-wide analysis
Drought stress
url https://doi.org/10.1186/s12870-025-06606-7
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