Microrchidia OsMORC6 Positively Regulates Cadmium Tolerance and Uptake by Mediating DNA Methylation in Rice

Abstract Rice (Oryza sativa) is an extremely important global food crop. However, cadmium (Cd) contamination in paddy fields poses a serious threat to human health worldwide. To generate low-Cd or Cd-free rice germplasms, it is essential to understand the molecular mechanisms involved in Cd toleranc...

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Main Authors: Jingai Tan, Muhammad Fahad, Lantian Zhang, Liang Wu, Xia Wu
Format: Article
Language:English
Published: SpringerOpen 2025-04-01
Series:Rice
Subjects:
Online Access:https://doi.org/10.1186/s12284-025-00785-5
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author Jingai Tan
Muhammad Fahad
Lantian Zhang
Liang Wu
Xia Wu
author_facet Jingai Tan
Muhammad Fahad
Lantian Zhang
Liang Wu
Xia Wu
author_sort Jingai Tan
collection DOAJ
description Abstract Rice (Oryza sativa) is an extremely important global food crop. However, cadmium (Cd) contamination in paddy fields poses a serious threat to human health worldwide. To generate low-Cd or Cd-free rice germplasms, it is essential to understand the molecular mechanisms involved in Cd tolerance, uptake, and translocation from soil to plant. In this study, we identify three Microrchidia proteins, OsMORC6a, OsMORC6b, and OsMORC6c, that regulate Cd tolerance and accumulation, although they do not alter the translocation of Cd from roots to shoots. Knockout of all three genes results in reducing Cd accumulation and increasing sensitivity to Cd stress. Furthermore, transcriptome analysis reveals 1,127 differentially expressed genes (DEGs) in the morc6abc mutants, which are significantly enriched in ‘plant-type cell wall’ and ‘oxidoreductase activity’ pathways. Through an integrating DNA methylome and transcriptome data, we identify 247 hyper-DMR-associated DEGs and 325 hypo-DMR-associated DEGs in morc6abc mutants. Gene Ontology (Go) enrichment analysis reveals that OsMORC6 proteins positively regulate Cd tolerance and uptake by mediating DNA methylation, which regulates the proper expression of genes related to plant cell wall and oxidative stress under Cd stress. Taken together, our findings reveal novel genes that mediate Cd tolerance and accumulation by affecting DNA methylation, offering valuable resource for breeding low-Cd or Cd-free rice germplasms.
format Article
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issn 1939-8425
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language English
publishDate 2025-04-01
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spelling doaj-art-9b0cdf5bb92f4f9d84e583818a12d5192025-08-20T02:17:10ZengSpringerOpenRice1939-84251939-84332025-04-0118111710.1186/s12284-025-00785-5Microrchidia OsMORC6 Positively Regulates Cadmium Tolerance and Uptake by Mediating DNA Methylation in RiceJingai Tan0Muhammad Fahad1Lantian Zhang2Liang Wu3Xia Wu4Hainan Yazhou Bay Seed Laboratory, Hainan Institute, Zhejiang UniversityHainan Yazhou Bay Seed Laboratory, Hainan Institute, Zhejiang UniversityHainan Yazhou Bay Seed Laboratory, Hainan Institute, Zhejiang UniversityHainan Yazhou Bay Seed Laboratory, Hainan Institute, Zhejiang UniversityHainan Yazhou Bay Seed Laboratory, Hainan Institute, Zhejiang UniversityAbstract Rice (Oryza sativa) is an extremely important global food crop. However, cadmium (Cd) contamination in paddy fields poses a serious threat to human health worldwide. To generate low-Cd or Cd-free rice germplasms, it is essential to understand the molecular mechanisms involved in Cd tolerance, uptake, and translocation from soil to plant. In this study, we identify three Microrchidia proteins, OsMORC6a, OsMORC6b, and OsMORC6c, that regulate Cd tolerance and accumulation, although they do not alter the translocation of Cd from roots to shoots. Knockout of all three genes results in reducing Cd accumulation and increasing sensitivity to Cd stress. Furthermore, transcriptome analysis reveals 1,127 differentially expressed genes (DEGs) in the morc6abc mutants, which are significantly enriched in ‘plant-type cell wall’ and ‘oxidoreductase activity’ pathways. Through an integrating DNA methylome and transcriptome data, we identify 247 hyper-DMR-associated DEGs and 325 hypo-DMR-associated DEGs in morc6abc mutants. Gene Ontology (Go) enrichment analysis reveals that OsMORC6 proteins positively regulate Cd tolerance and uptake by mediating DNA methylation, which regulates the proper expression of genes related to plant cell wall and oxidative stress under Cd stress. Taken together, our findings reveal novel genes that mediate Cd tolerance and accumulation by affecting DNA methylation, offering valuable resource for breeding low-Cd or Cd-free rice germplasms.https://doi.org/10.1186/s12284-025-00785-5CdOsMORC6aOsMORC6bOsMORC6cToleranceAccumulation
spellingShingle Jingai Tan
Muhammad Fahad
Lantian Zhang
Liang Wu
Xia Wu
Microrchidia OsMORC6 Positively Regulates Cadmium Tolerance and Uptake by Mediating DNA Methylation in Rice
Rice
Cd
OsMORC6a
OsMORC6b
OsMORC6c
Tolerance
Accumulation
title Microrchidia OsMORC6 Positively Regulates Cadmium Tolerance and Uptake by Mediating DNA Methylation in Rice
title_full Microrchidia OsMORC6 Positively Regulates Cadmium Tolerance and Uptake by Mediating DNA Methylation in Rice
title_fullStr Microrchidia OsMORC6 Positively Regulates Cadmium Tolerance and Uptake by Mediating DNA Methylation in Rice
title_full_unstemmed Microrchidia OsMORC6 Positively Regulates Cadmium Tolerance and Uptake by Mediating DNA Methylation in Rice
title_short Microrchidia OsMORC6 Positively Regulates Cadmium Tolerance and Uptake by Mediating DNA Methylation in Rice
title_sort microrchidia osmorc6 positively regulates cadmium tolerance and uptake by mediating dna methylation in rice
topic Cd
OsMORC6a
OsMORC6b
OsMORC6c
Tolerance
Accumulation
url https://doi.org/10.1186/s12284-025-00785-5
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