Dipole Potential of Monolayers with Biologically Relevant Lipid Compositions

The membrane dipole potential that arises from the interfacial water and constitutive dipolar groups of lipid molecules modulates the interaction of amphiphiles and proteins with membranes. Consequently, its determination for lipid mixtures resembling the existing diversity in biological membranes i...

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Main Authors: Renato M. S. Cardoso, Fabiana Lairion, Edgardo Anibal Disalvo, Luís M. S. Loura, Maria João Moreno
Format: Article
Language:English
Published: MDPI AG 2024-12-01
Series:Molecules
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Online Access:https://www.mdpi.com/1420-3049/29/24/5843
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author Renato M. S. Cardoso
Fabiana Lairion
Edgardo Anibal Disalvo
Luís M. S. Loura
Maria João Moreno
author_facet Renato M. S. Cardoso
Fabiana Lairion
Edgardo Anibal Disalvo
Luís M. S. Loura
Maria João Moreno
author_sort Renato M. S. Cardoso
collection DOAJ
description The membrane dipole potential that arises from the interfacial water and constitutive dipolar groups of lipid molecules modulates the interaction of amphiphiles and proteins with membranes. Consequently, its determination for lipid mixtures resembling the existing diversity in biological membranes is very relevant. In this work, the dipole potentials of monolayers, formed at the air–water interface, from pure or mixed lipids (1-palmitoyl-2-oleoyl-<i>sn</i>-glycero-3-phosphocholine (POPC), 1-palmitoyl-2-oleoyl-<i>sn</i>-glycero-3-phosphoethanolamine (POPE), 1-palmitoyl-2-oleoyl-<i>sn</i>-glycero-3-phosphatidyserine (POPS), sphingomyelin (SpM) and cholesterol) were measured and correlated with the mean area per lipid. The results showed that, as previously observed, cholesterol increases the dipole potential in correspondence with the decrease in the average area per lipid. At the small mole fractions encountered in biomembranes, the presence of the negatively charged lipid POPS increases the dipole potentials of monolayers despite inducing an increase in the average area per lipid. Additionally, the inclusion of POPE in POPC:cholesterol monolayers disrupts the area condensation induced by cholesterol while increasing the membrane dipole moment, leading to a small reduction in the dipole potential. This trend is reinforced for the quaternary POPC:cholesterol:POPE:POPS 4:3:2:1 system, which mimics the inner leaflets of eukaryotic plasma membranes. In agreement with previous works, the replacement of phosphocholine lipids with sphingomyelin leads to a decrease in the dipole potential. Together, this results in a lower dipole potential for the SpM-enriched outer leaflet, generating a non-zero transbilayer dipole potential in the asymmetric plasma membranes of eukaryotic cells.
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spelling doaj-art-9aa198b9609e4b07995d2f9fc8b7e55e2025-08-20T02:01:28ZengMDPI AGMolecules1420-30492024-12-012924584310.3390/molecules29245843Dipole Potential of Monolayers with Biologically Relevant Lipid CompositionsRenato M. S. Cardoso0Fabiana Lairion1Edgardo Anibal Disalvo2Luís M. S. Loura3Maria João Moreno4Coimbra Chemistry Center, Institute of Molecular Sciences (CQC-IMS), University of Coimbra, 3004-535 Coimbra, PortugalInstitute of Biochemistry and Molecular Medicine Prof. Alberto Boveris (IBIMOL), University of Buenos Aires and National Council for Scientific and Technical Research (CONICET), Buenos Aires 1113, ArgentinaApplied Biophysics and Food Research Center (CIBAAL), National University of Santiago del Estero and National Council for Scientific and Technical Research (CONICET), Santiago del Estero 4206, ArgentinaCoimbra Chemistry Center, Institute of Molecular Sciences (CQC-IMS), University of Coimbra, 3004-535 Coimbra, PortugalCoimbra Chemistry Center, Institute of Molecular Sciences (CQC-IMS), University of Coimbra, 3004-535 Coimbra, PortugalThe membrane dipole potential that arises from the interfacial water and constitutive dipolar groups of lipid molecules modulates the interaction of amphiphiles and proteins with membranes. Consequently, its determination for lipid mixtures resembling the existing diversity in biological membranes is very relevant. In this work, the dipole potentials of monolayers, formed at the air–water interface, from pure or mixed lipids (1-palmitoyl-2-oleoyl-<i>sn</i>-glycero-3-phosphocholine (POPC), 1-palmitoyl-2-oleoyl-<i>sn</i>-glycero-3-phosphoethanolamine (POPE), 1-palmitoyl-2-oleoyl-<i>sn</i>-glycero-3-phosphatidyserine (POPS), sphingomyelin (SpM) and cholesterol) were measured and correlated with the mean area per lipid. The results showed that, as previously observed, cholesterol increases the dipole potential in correspondence with the decrease in the average area per lipid. At the small mole fractions encountered in biomembranes, the presence of the negatively charged lipid POPS increases the dipole potentials of monolayers despite inducing an increase in the average area per lipid. Additionally, the inclusion of POPE in POPC:cholesterol monolayers disrupts the area condensation induced by cholesterol while increasing the membrane dipole moment, leading to a small reduction in the dipole potential. This trend is reinforced for the quaternary POPC:cholesterol:POPE:POPS 4:3:2:1 system, which mimics the inner leaflets of eukaryotic plasma membranes. In agreement with previous works, the replacement of phosphocholine lipids with sphingomyelin leads to a decrease in the dipole potential. Together, this results in a lower dipole potential for the SpM-enriched outer leaflet, generating a non-zero transbilayer dipole potential in the asymmetric plasma membranes of eukaryotic cells.https://www.mdpi.com/1420-3049/29/24/5843transmembrane potentialcholesterolPOPCPOPEPOPSsurface pressure
spellingShingle Renato M. S. Cardoso
Fabiana Lairion
Edgardo Anibal Disalvo
Luís M. S. Loura
Maria João Moreno
Dipole Potential of Monolayers with Biologically Relevant Lipid Compositions
Molecules
transmembrane potential
cholesterol
POPC
POPE
POPS
surface pressure
title Dipole Potential of Monolayers with Biologically Relevant Lipid Compositions
title_full Dipole Potential of Monolayers with Biologically Relevant Lipid Compositions
title_fullStr Dipole Potential of Monolayers with Biologically Relevant Lipid Compositions
title_full_unstemmed Dipole Potential of Monolayers with Biologically Relevant Lipid Compositions
title_short Dipole Potential of Monolayers with Biologically Relevant Lipid Compositions
title_sort dipole potential of monolayers with biologically relevant lipid compositions
topic transmembrane potential
cholesterol
POPC
POPE
POPS
surface pressure
url https://www.mdpi.com/1420-3049/29/24/5843
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