Vascular bundle cell-specific expression of a phosphate transporter improves phosphate use efficiency of transgenic Arabidopsis without detrimental effects

Abstract Constitutive overexpression of phosphate (Pi) transporter family 1 often results in the accumulation of toxic levels of Pi, which causes growth retardation in plants. In contrast, we had previously reported that root epidermis-specific overexpression of the phosphate transporter TaPT2 in Ar...

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Main Authors: Yuichi Tada, Aoi Shimizu
Format: Article
Language:English
Published: Nature Portfolio 2024-11-01
Series:Scientific Reports
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Online Access:https://doi.org/10.1038/s41598-024-78500-5
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author Yuichi Tada
Aoi Shimizu
author_facet Yuichi Tada
Aoi Shimizu
author_sort Yuichi Tada
collection DOAJ
description Abstract Constitutive overexpression of phosphate (Pi) transporter family 1 often results in the accumulation of toxic levels of Pi, which causes growth retardation in plants. In contrast, we had previously reported that root epidermis-specific overexpression of the phosphate transporter TaPT2 in Arabidopsis leads to improved growth and Pi use efficiency. In the present study, we used promoters AtHKT1;1 and SKOR, which are predominantly expressed in the vascular bundle tissues, to overexpress TaPT2. Transgenic lines exhibited increased shoot growth compared to wild type plants under normal- and low-Pi conditions, along with elevated root Pi and total P content, and higher xylem sap Pi concentration, specifically under low-Pi conditions. This was attributed to moderate Pi accumulation in the xylem parenchyma cells, enhancing the Pi uploading capacity to the xylem. SKOR-TaPT2, however, did not complement pho1 mutant, which was defective in uploading Pi to the xylem. The transcriptional levels of VPT1 and VPT3, which are responsible for transporting excess Pi into a vacuole, were upregulated in SKOR promoter lines under normal-Pi conditions. Our results suggested that root vascular bundle-specific expression of TaPT2 is another promising strategy for increasing biomass production, Pi uptake, and Pi use efficiency while preventing growth retardation in transgenic plants.
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spelling doaj-art-9a7e5ee57e074ae1af1f095fb31524f02025-08-20T02:50:04ZengNature PortfolioScientific Reports2045-23222024-11-0114111010.1038/s41598-024-78500-5Vascular bundle cell-specific expression of a phosphate transporter improves phosphate use efficiency of transgenic Arabidopsis without detrimental effectsYuichi Tada0Aoi Shimizu1School of Bioscience and Biotechnology, Tokyo University of TechnologyGraduate School of Bionics, Tokyo University of TechnologyAbstract Constitutive overexpression of phosphate (Pi) transporter family 1 often results in the accumulation of toxic levels of Pi, which causes growth retardation in plants. In contrast, we had previously reported that root epidermis-specific overexpression of the phosphate transporter TaPT2 in Arabidopsis leads to improved growth and Pi use efficiency. In the present study, we used promoters AtHKT1;1 and SKOR, which are predominantly expressed in the vascular bundle tissues, to overexpress TaPT2. Transgenic lines exhibited increased shoot growth compared to wild type plants under normal- and low-Pi conditions, along with elevated root Pi and total P content, and higher xylem sap Pi concentration, specifically under low-Pi conditions. This was attributed to moderate Pi accumulation in the xylem parenchyma cells, enhancing the Pi uploading capacity to the xylem. SKOR-TaPT2, however, did not complement pho1 mutant, which was defective in uploading Pi to the xylem. The transcriptional levels of VPT1 and VPT3, which are responsible for transporting excess Pi into a vacuole, were upregulated in SKOR promoter lines under normal-Pi conditions. Our results suggested that root vascular bundle-specific expression of TaPT2 is another promising strategy for increasing biomass production, Pi uptake, and Pi use efficiency while preventing growth retardation in transgenic plants.https://doi.org/10.1038/s41598-024-78500-5Growth promotionPhosphate transporterPi acquisitionPi use efficiencyVascular bundle cell-specific expression
spellingShingle Yuichi Tada
Aoi Shimizu
Vascular bundle cell-specific expression of a phosphate transporter improves phosphate use efficiency of transgenic Arabidopsis without detrimental effects
Scientific Reports
Growth promotion
Phosphate transporter
Pi acquisition
Pi use efficiency
Vascular bundle cell-specific expression
title Vascular bundle cell-specific expression of a phosphate transporter improves phosphate use efficiency of transgenic Arabidopsis without detrimental effects
title_full Vascular bundle cell-specific expression of a phosphate transporter improves phosphate use efficiency of transgenic Arabidopsis without detrimental effects
title_fullStr Vascular bundle cell-specific expression of a phosphate transporter improves phosphate use efficiency of transgenic Arabidopsis without detrimental effects
title_full_unstemmed Vascular bundle cell-specific expression of a phosphate transporter improves phosphate use efficiency of transgenic Arabidopsis without detrimental effects
title_short Vascular bundle cell-specific expression of a phosphate transporter improves phosphate use efficiency of transgenic Arabidopsis without detrimental effects
title_sort vascular bundle cell specific expression of a phosphate transporter improves phosphate use efficiency of transgenic arabidopsis without detrimental effects
topic Growth promotion
Phosphate transporter
Pi acquisition
Pi use efficiency
Vascular bundle cell-specific expression
url https://doi.org/10.1038/s41598-024-78500-5
work_keys_str_mv AT yuichitada vascularbundlecellspecificexpressionofaphosphatetransporterimprovesphosphateuseefficiencyoftransgenicarabidopsiswithoutdetrimentaleffects
AT aoishimizu vascularbundlecellspecificexpressionofaphosphatetransporterimprovesphosphateuseefficiencyoftransgenicarabidopsiswithoutdetrimentaleffects