Arabidopsis root apical meristem adaptation to an osmotic gradient condition: an integrated approach from cell expansion to gene expression

Climate change triggers abiotic stress, such as drought and high salinity, that can cause osmotic stress. Water availability can limit plant growth, and the root tip tissues initially sense it. Most experiments destined to understand root growth adaptation to osmotic stress apply homogeneous high os...

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Main Authors: Selene Píriz-Pezzutto, Mauro Martínez-Moré, María Martha Sainz, Omar Borsani, Mariana Sotelo-Silveira
Format: Article
Language:English
Published: Frontiers Media S.A. 2024-11-01
Series:Frontiers in Plant Science
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Online Access:https://www.frontiersin.org/articles/10.3389/fpls.2024.1465219/full
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author Selene Píriz-Pezzutto
Mauro Martínez-Moré
María Martha Sainz
Omar Borsani
Mariana Sotelo-Silveira
author_facet Selene Píriz-Pezzutto
Mauro Martínez-Moré
María Martha Sainz
Omar Borsani
Mariana Sotelo-Silveira
author_sort Selene Píriz-Pezzutto
collection DOAJ
description Climate change triggers abiotic stress, such as drought and high salinity, that can cause osmotic stress. Water availability can limit plant growth, and the root tip tissues initially sense it. Most experiments destined to understand root growth adaptation to osmotic stress apply homogeneous high osmotic potentials (osmotic shock) to shoots and roots. However, this treatment does not represent natural field conditions where a root may encounter increasing osmotic potentials while exploring the soil. Osmotic shock severely reduces root growth rate, decreasing cell division in the proximal meristem and reducing mature cell length. In this work, we developed an in vitro osmotic gradient experimental system with increasing osmotic potentials. The system generates a controlled osmotic gradient in the root growth zone while exposing the aerial tissues to control conditions. The osmotic gradient system allowed Arabidopsis seedlings of Col-0 and ttl1 mutant (affected in the gene TETRATRICOPEPTIDE THIOREDOXIN-LIKE 1 (TTL1)) to sustain proper root growth for 25 days, reaching osmotic potentials of -1.2 MPa. We demonstrated that roots of seedlings grown in the osmotic gradient sustain a higher root growth rate than those that were grown under a homogeneous high osmotic potential. Furthermore, we found out that the expression of some genes is modified in the roots grown in the osmotic gradient compared to those grown in osmotic shock. Our data indicate that using an osmotic gradient can improve our understanding of how plants respond to osmotic stress and help find new genes to improve plant field performance.
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spelling doaj-art-9654cc2bfbf34ea8a6b20aaffe9551a62025-08-20T02:13:01ZengFrontiers Media S.A.Frontiers in Plant Science1664-462X2024-11-011510.3389/fpls.2024.14652191465219Arabidopsis root apical meristem adaptation to an osmotic gradient condition: an integrated approach from cell expansion to gene expressionSelene Píriz-PezzuttoMauro Martínez-MoréMaría Martha SainzOmar BorsaniMariana Sotelo-SilveiraClimate change triggers abiotic stress, such as drought and high salinity, that can cause osmotic stress. Water availability can limit plant growth, and the root tip tissues initially sense it. Most experiments destined to understand root growth adaptation to osmotic stress apply homogeneous high osmotic potentials (osmotic shock) to shoots and roots. However, this treatment does not represent natural field conditions where a root may encounter increasing osmotic potentials while exploring the soil. Osmotic shock severely reduces root growth rate, decreasing cell division in the proximal meristem and reducing mature cell length. In this work, we developed an in vitro osmotic gradient experimental system with increasing osmotic potentials. The system generates a controlled osmotic gradient in the root growth zone while exposing the aerial tissues to control conditions. The osmotic gradient system allowed Arabidopsis seedlings of Col-0 and ttl1 mutant (affected in the gene TETRATRICOPEPTIDE THIOREDOXIN-LIKE 1 (TTL1)) to sustain proper root growth for 25 days, reaching osmotic potentials of -1.2 MPa. We demonstrated that roots of seedlings grown in the osmotic gradient sustain a higher root growth rate than those that were grown under a homogeneous high osmotic potential. Furthermore, we found out that the expression of some genes is modified in the roots grown in the osmotic gradient compared to those grown in osmotic shock. Our data indicate that using an osmotic gradient can improve our understanding of how plants respond to osmotic stress and help find new genes to improve plant field performance.https://www.frontiersin.org/articles/10.3389/fpls.2024.1465219/fullosmotic stressgradientroot growthArabidopsisTTL1
spellingShingle Selene Píriz-Pezzutto
Mauro Martínez-Moré
María Martha Sainz
Omar Borsani
Mariana Sotelo-Silveira
Arabidopsis root apical meristem adaptation to an osmotic gradient condition: an integrated approach from cell expansion to gene expression
Frontiers in Plant Science
osmotic stress
gradient
root growth
Arabidopsis
TTL1
title Arabidopsis root apical meristem adaptation to an osmotic gradient condition: an integrated approach from cell expansion to gene expression
title_full Arabidopsis root apical meristem adaptation to an osmotic gradient condition: an integrated approach from cell expansion to gene expression
title_fullStr Arabidopsis root apical meristem adaptation to an osmotic gradient condition: an integrated approach from cell expansion to gene expression
title_full_unstemmed Arabidopsis root apical meristem adaptation to an osmotic gradient condition: an integrated approach from cell expansion to gene expression
title_short Arabidopsis root apical meristem adaptation to an osmotic gradient condition: an integrated approach from cell expansion to gene expression
title_sort arabidopsis root apical meristem adaptation to an osmotic gradient condition an integrated approach from cell expansion to gene expression
topic osmotic stress
gradient
root growth
Arabidopsis
TTL1
url https://www.frontiersin.org/articles/10.3389/fpls.2024.1465219/full
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AT mariamarthasainz arabidopsisrootapicalmeristemadaptationtoanosmoticgradientconditionanintegratedapproachfromcellexpansiontogeneexpression
AT omarborsani arabidopsisrootapicalmeristemadaptationtoanosmoticgradientconditionanintegratedapproachfromcellexpansiontogeneexpression
AT marianasotelosilveira arabidopsisrootapicalmeristemadaptationtoanosmoticgradientconditionanintegratedapproachfromcellexpansiontogeneexpression