Knocking down the neuronal lactate transporter MCT2 in the arcuate nucleus of female rats increases food intake and body weight

Abstract In the arcuate nucleus of the hypothalamus, tanycyte-neuron interactions regulate glucose homeostasis and feeding behavior. Previously, we reported that monocarboxylate transporters (MCT) 1 and 4 are localized in tanycytes, whereas MCT2 is present in arcuate nucleus neurons, including orexi...

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Main Authors: Alanis Coca, Sergio López, Patricio Órdenes, Vania Sepúlveda, Diego Cuevas, Andrés Villarroel, Javiera Álvarez-Indo, Patricia V. Burgos, Estefanía Tarifeño, Roberto Elizondo-Vega, María A García-Robles
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Language:English
Published: Nature Portfolio 2025-03-01
Series:Scientific Reports
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Online Access:https://doi.org/10.1038/s41598-025-90513-2
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author Alanis Coca
Sergio López
Patricio Órdenes
Vania Sepúlveda
Diego Cuevas
Andrés Villarroel
Javiera Álvarez-Indo
Patricia V. Burgos
Estefanía Tarifeño
Roberto Elizondo-Vega
María A García-Robles
author_facet Alanis Coca
Sergio López
Patricio Órdenes
Vania Sepúlveda
Diego Cuevas
Andrés Villarroel
Javiera Álvarez-Indo
Patricia V. Burgos
Estefanía Tarifeño
Roberto Elizondo-Vega
María A García-Robles
author_sort Alanis Coca
collection DOAJ
description Abstract In the arcuate nucleus of the hypothalamus, tanycyte-neuron interactions regulate glucose homeostasis and feeding behavior. Previously, we reported that monocarboxylate transporters (MCT) 1 and 4 are localized in tanycytes, whereas MCT2 is present in arcuate nucleus neurons, including orexigenic and anorexigenic neurons (POMC). MCT1 and MCT4 inhibition impacts feeding behavior, suggesting that monocarboxylate transfer between tanycytes and neurons influences food intake. Electrophysiological studies have shown that POMC neurons respond to lactate through transport and indirect signaling using astrocytic hydroxycarboxylic acid receptor 1. To investigate the role of MCT2 further, we generated MCT2 knockdown rats and analyzed their feeding behavior. Female Sprague-Dawley rats received bilateral injections in the arcuate nucleus with an adeno-associated virus (AAV) carrying a specific short hairpin RNA to inhibit MCT2 expression, thereby generating neuronal MCT2 knockdown rats. Knockdown efficiency in rat hypothalamic tissue was assessed using real-time PCR, Western Blot, and immunohistochemistry. The acute effect on feeding behavior was evaluated following 24 h of fasting, followed by 24 h of refeeding. In MCT2-knockdown rats, we observed additional inhibition of MCT1, suggesting a potential glial response to increased parenchymal lactate levels. Both macrostructure and microstructure of feeding were evaluated in MCT2-knockdown rats and compared to control AAV-injected rats. MCT2 knockdown led to a significant increase in macrostructural parameters, such as food intake and body weight. These findings underscore the importance of lactate transfer as a mechanism in tanycyte-neuron communication mediated by monocarboxylates.
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spelling doaj-art-9295b4b722cb4c9997b95e7a99cf463c2025-08-20T03:05:56ZengNature PortfolioScientific Reports2045-23222025-03-0115111310.1038/s41598-025-90513-2Knocking down the neuronal lactate transporter MCT2 in the arcuate nucleus of female rats increases food intake and body weightAlanis Coca0Sergio López1Patricio Órdenes2Vania Sepúlveda3Diego Cuevas4Andrés Villarroel5Javiera Álvarez-Indo6Patricia V. Burgos7Estefanía Tarifeño8Roberto Elizondo-Vega9María A García-Robles10Laboratorio de Biología Celular, Departamento de Biología Celular, Facultad de Ciencias Biológicas, Universidad de ConcepciónLaboratorio de Biología Celular, Departamento de Biología Celular, Facultad de Ciencias Biológicas, Universidad de ConcepciónLaboratorio de Neuroinmunología, Facultad de Medicina y Ciencia, Universidad San Sebastián, Sede ConcepciónLaboratorio de Biología Celular, Departamento de Biología Celular, Facultad de Ciencias Biológicas, Universidad de ConcepciónLaboratorio de Biología Celular, Departamento de Biología Celular, Facultad de Ciencias Biológicas, Universidad de ConcepciónLaboratorio de Biología Celular, Departamento de Biología Celular, Facultad de Ciencias Biológicas, Universidad de ConcepciónCentro de Biología Celular y Biomedicina, Facultad de Medicina y Ciencia, Universidad San SebastiánCentro de Biología Celular y Biomedicina, Facultad de Medicina y Ciencia, Universidad San SebastiánLaboratorio de Expresión y Regulación Génica, Departamento de Bioquímica y Biología Molecular, Facultad de Ciencias Biológicas, Universidad de ConcepciónLaboratorio de Biología Celular, Departamento de Biología Celular, Facultad de Ciencias Biológicas, Universidad de ConcepciónLaboratorio de Biología Celular, Departamento de Biología Celular, Facultad de Ciencias Biológicas, Universidad de ConcepciónAbstract In the arcuate nucleus of the hypothalamus, tanycyte-neuron interactions regulate glucose homeostasis and feeding behavior. Previously, we reported that monocarboxylate transporters (MCT) 1 and 4 are localized in tanycytes, whereas MCT2 is present in arcuate nucleus neurons, including orexigenic and anorexigenic neurons (POMC). MCT1 and MCT4 inhibition impacts feeding behavior, suggesting that monocarboxylate transfer between tanycytes and neurons influences food intake. Electrophysiological studies have shown that POMC neurons respond to lactate through transport and indirect signaling using astrocytic hydroxycarboxylic acid receptor 1. To investigate the role of MCT2 further, we generated MCT2 knockdown rats and analyzed their feeding behavior. Female Sprague-Dawley rats received bilateral injections in the arcuate nucleus with an adeno-associated virus (AAV) carrying a specific short hairpin RNA to inhibit MCT2 expression, thereby generating neuronal MCT2 knockdown rats. Knockdown efficiency in rat hypothalamic tissue was assessed using real-time PCR, Western Blot, and immunohistochemistry. The acute effect on feeding behavior was evaluated following 24 h of fasting, followed by 24 h of refeeding. In MCT2-knockdown rats, we observed additional inhibition of MCT1, suggesting a potential glial response to increased parenchymal lactate levels. Both macrostructure and microstructure of feeding were evaluated in MCT2-knockdown rats and compared to control AAV-injected rats. MCT2 knockdown led to a significant increase in macrostructural parameters, such as food intake and body weight. These findings underscore the importance of lactate transfer as a mechanism in tanycyte-neuron communication mediated by monocarboxylates.https://doi.org/10.1038/s41598-025-90513-2MCT2Arcuate nucleusFeeding behaviorRatsSatiation
spellingShingle Alanis Coca
Sergio López
Patricio Órdenes
Vania Sepúlveda
Diego Cuevas
Andrés Villarroel
Javiera Álvarez-Indo
Patricia V. Burgos
Estefanía Tarifeño
Roberto Elizondo-Vega
María A García-Robles
Knocking down the neuronal lactate transporter MCT2 in the arcuate nucleus of female rats increases food intake and body weight
Scientific Reports
MCT2
Arcuate nucleus
Feeding behavior
Rats
Satiation
title Knocking down the neuronal lactate transporter MCT2 in the arcuate nucleus of female rats increases food intake and body weight
title_full Knocking down the neuronal lactate transporter MCT2 in the arcuate nucleus of female rats increases food intake and body weight
title_fullStr Knocking down the neuronal lactate transporter MCT2 in the arcuate nucleus of female rats increases food intake and body weight
title_full_unstemmed Knocking down the neuronal lactate transporter MCT2 in the arcuate nucleus of female rats increases food intake and body weight
title_short Knocking down the neuronal lactate transporter MCT2 in the arcuate nucleus of female rats increases food intake and body weight
title_sort knocking down the neuronal lactate transporter mct2 in the arcuate nucleus of female rats increases food intake and body weight
topic MCT2
Arcuate nucleus
Feeding behavior
Rats
Satiation
url https://doi.org/10.1038/s41598-025-90513-2
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