Thinning intensity influences the C:N:P stoichiometry in forest ecosystems: A global synthesis

Forest thinning potentially alters carbon (C), nitrogen (N), and phosphorus (P) cycles, thereby affecting their C:N:P stoichiometry as well as the key ecosystem services they support. Despite the fact that numerous individual studies and a few meta-analyses have been conducted to examine thinning ef...

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Main Authors: Chongwei Fan, Guiyao Zhou, Hongyang Chen, Zhenggang Du, Ruiqiang Liu, Yanghui He, Changjiang Huang, Shuying Qiu, Yimin Zhu, Jie Li, Xuhui Zhou
Format: Article
Language:English
Published: Elsevier 2025-08-01
Series:Geoderma
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Online Access:http://www.sciencedirect.com/science/article/pii/S0016706125002769
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author Chongwei Fan
Guiyao Zhou
Hongyang Chen
Zhenggang Du
Ruiqiang Liu
Yanghui He
Changjiang Huang
Shuying Qiu
Yimin Zhu
Jie Li
Xuhui Zhou
author_facet Chongwei Fan
Guiyao Zhou
Hongyang Chen
Zhenggang Du
Ruiqiang Liu
Yanghui He
Changjiang Huang
Shuying Qiu
Yimin Zhu
Jie Li
Xuhui Zhou
author_sort Chongwei Fan
collection DOAJ
description Forest thinning potentially alters carbon (C), nitrogen (N), and phosphorus (P) cycles, thereby affecting their C:N:P stoichiometry as well as the key ecosystem services they support. Despite the fact that numerous individual studies and a few meta-analyses have been conducted to examine thinning effects on ecosystem C and N cycles, how forest thinning, especially its intensity, affects the C:N:P stoichiometry of plants, soil, and microbes remains poorly known. Here, we carried out a global meta-analysis of 779 paired observations from 136 peer-reviewed articles to assess effects of forest thinning on C:N:P stoichiometry of plants, soil, and microbes in global forests. Our results showed that, on average, forest thinning significantly increased soil C and P pools by 5.0% and 11.1%, and microbial biomass C, N, and, P pools by 9.3%, 12.0%, and 38.4%, respectively. In contrast, forest thinning decreased plant P pool, soil C:P and N:P ratios, and microbial C:N and N:P ratios. More importantly, the effects of forest thinning on C:N:P stoichiometry largely varied with forest thinning intensity. Specifically, heavy thinning significantly decreased plant C:N ratios and soil N:P ratios, whereas light and moderate thinning had positive or insignificant effects on C:N:P stoichiometry. Thinning-induced changes in the C:N, C:P, and N:P ratios were positively correlated with mean annual temperature (MAT) and mean annual precipitation (MAP), but negatively with post-thinning duration and stand age. These findings highlight the impact of thinning intensity on the connection between above- and belowground processes, which may decouple the biogeochemical cycles of C, N, and P in forest ecosystems. Future forest management could implement an appropriate forest thinning intensity to ensure ecosystem functionality and maintain the balance and stability of ecosystem elements.
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spelling doaj-art-9133fefee80c4d77a0b2bd06f4bd799b2025-08-20T03:02:30ZengElsevierGeoderma1872-62592025-08-0146011743510.1016/j.geoderma.2025.117435Thinning intensity influences the C:N:P stoichiometry in forest ecosystems: A global synthesisChongwei Fan0Guiyao Zhou1Hongyang Chen2Zhenggang Du3Ruiqiang Liu4Yanghui He5Changjiang Huang6Shuying Qiu7Yimin Zhu8Jie Li9Xuhui Zhou10Institute of Carbon Neutrality, Key Laboratory of Sustainable Forest Ecosystem Management-Ministry of Education, School of Ecology, Northeast Forestry University, Harbin 150040, China; School of Future Technology, Northeast Forestry University, Harbin 150040, ChinaLaboratorio de Biodiversidad y Funcionamiento Ecosistémico. Instituto de Recursos Naturales y Agrobiología de Sevilla (IRNAS), CSIC, Av. Reina Mercedes 10, E-41012 Sevilla, SpainInstitute of Carbon Neutrality, Key Laboratory of Sustainable Forest Ecosystem Management-Ministry of Education, School of Ecology, Northeast Forestry University, Harbin 150040, China; School of Future Technology, Northeast Forestry University, Harbin 150040, ChinaInstitute of Carbon Neutrality, Key Laboratory of Sustainable Forest Ecosystem Management-Ministry of Education, School of Ecology, Northeast Forestry University, Harbin 150040, ChinaInstitute of Carbon Neutrality, Key Laboratory of Sustainable Forest Ecosystem Management-Ministry of Education, School of Ecology, Northeast Forestry University, Harbin 150040, ChinaInstitute of Carbon Neutrality, Key Laboratory of Sustainable Forest Ecosystem Management-Ministry of Education, School of Ecology, Northeast Forestry University, Harbin 150040, China; School of Future Technology, Northeast Forestry University, Harbin 150040, ChinaInstitute of Carbon Neutrality, Key Laboratory of Sustainable Forest Ecosystem Management-Ministry of Education, School of Ecology, Northeast Forestry University, Harbin 150040, ChinaInstitute of Carbon Neutrality, Key Laboratory of Sustainable Forest Ecosystem Management-Ministry of Education, School of Ecology, Northeast Forestry University, Harbin 150040, ChinaInstitute of Carbon Neutrality, Key Laboratory of Sustainable Forest Ecosystem Management-Ministry of Education, School of Ecology, Northeast Forestry University, Harbin 150040, ChinaInstitute of Carbon Neutrality, Key Laboratory of Sustainable Forest Ecosystem Management-Ministry of Education, School of Ecology, Northeast Forestry University, Harbin 150040, ChinaInstitute of Carbon Neutrality, Key Laboratory of Sustainable Forest Ecosystem Management-Ministry of Education, School of Ecology, Northeast Forestry University, Harbin 150040, China; School of Future Technology, Northeast Forestry University, Harbin 150040, China; Corresponding author at: School of Ecology, Northeast Forestry University, 26 Hexing Road, Harbin 150040, China.Forest thinning potentially alters carbon (C), nitrogen (N), and phosphorus (P) cycles, thereby affecting their C:N:P stoichiometry as well as the key ecosystem services they support. Despite the fact that numerous individual studies and a few meta-analyses have been conducted to examine thinning effects on ecosystem C and N cycles, how forest thinning, especially its intensity, affects the C:N:P stoichiometry of plants, soil, and microbes remains poorly known. Here, we carried out a global meta-analysis of 779 paired observations from 136 peer-reviewed articles to assess effects of forest thinning on C:N:P stoichiometry of plants, soil, and microbes in global forests. Our results showed that, on average, forest thinning significantly increased soil C and P pools by 5.0% and 11.1%, and microbial biomass C, N, and, P pools by 9.3%, 12.0%, and 38.4%, respectively. In contrast, forest thinning decreased plant P pool, soil C:P and N:P ratios, and microbial C:N and N:P ratios. More importantly, the effects of forest thinning on C:N:P stoichiometry largely varied with forest thinning intensity. Specifically, heavy thinning significantly decreased plant C:N ratios and soil N:P ratios, whereas light and moderate thinning had positive or insignificant effects on C:N:P stoichiometry. Thinning-induced changes in the C:N, C:P, and N:P ratios were positively correlated with mean annual temperature (MAT) and mean annual precipitation (MAP), but negatively with post-thinning duration and stand age. These findings highlight the impact of thinning intensity on the connection between above- and belowground processes, which may decouple the biogeochemical cycles of C, N, and P in forest ecosystems. Future forest management could implement an appropriate forest thinning intensity to ensure ecosystem functionality and maintain the balance and stability of ecosystem elements.http://www.sciencedirect.com/science/article/pii/S0016706125002769Forest managementThinning intensityC:N:P stoichiometryPlant-soil-microbeAbove- and belowground processes
spellingShingle Chongwei Fan
Guiyao Zhou
Hongyang Chen
Zhenggang Du
Ruiqiang Liu
Yanghui He
Changjiang Huang
Shuying Qiu
Yimin Zhu
Jie Li
Xuhui Zhou
Thinning intensity influences the C:N:P stoichiometry in forest ecosystems: A global synthesis
Geoderma
Forest management
Thinning intensity
C:N:P stoichiometry
Plant-soil-microbe
Above- and belowground processes
title Thinning intensity influences the C:N:P stoichiometry in forest ecosystems: A global synthesis
title_full Thinning intensity influences the C:N:P stoichiometry in forest ecosystems: A global synthesis
title_fullStr Thinning intensity influences the C:N:P stoichiometry in forest ecosystems: A global synthesis
title_full_unstemmed Thinning intensity influences the C:N:P stoichiometry in forest ecosystems: A global synthesis
title_short Thinning intensity influences the C:N:P stoichiometry in forest ecosystems: A global synthesis
title_sort thinning intensity influences the c n p stoichiometry in forest ecosystems a global synthesis
topic Forest management
Thinning intensity
C:N:P stoichiometry
Plant-soil-microbe
Above- and belowground processes
url http://www.sciencedirect.com/science/article/pii/S0016706125002769
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