Large-scale phylogenomic insights into the evolution of the Hymenochaetales

The Hymenochaetales is an order with most species as wood-inhabiting fungi that have high phylogenetic complexity and morphological diversity. Species in this order play important roles in forest ecosystems and include wood decomposers, pathogens, and those that form ectomycorrhizal associations. Ho...

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Main Authors: Heng Zhao, Fang Wu, Sundy Maurice, Igor N. Pavlov, Konstantin V. Krutovsky, Hong-Gao Liu, Yuan Yuan, Yu-Cheng Dai
Format: Article
Language:English
Published: Taylor & Francis Group 2025-04-01
Series:Mycology
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Online Access:https://www.tandfonline.com/doi/10.1080/21501203.2024.2391527
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author Heng Zhao
Fang Wu
Sundy Maurice
Igor N. Pavlov
Konstantin V. Krutovsky
Hong-Gao Liu
Yuan Yuan
Yu-Cheng Dai
author_facet Heng Zhao
Fang Wu
Sundy Maurice
Igor N. Pavlov
Konstantin V. Krutovsky
Hong-Gao Liu
Yuan Yuan
Yu-Cheng Dai
author_sort Heng Zhao
collection DOAJ
description The Hymenochaetales is an order with most species as wood-inhabiting fungi that have high phylogenetic complexity and morphological diversity. Species in this order play important roles in forest ecosystems and include wood decomposers, pathogens, and those that form ectomycorrhizal associations. However, we have limited knowledge of the patterns of large-scale evolutionary history of the order. In this study, using 171 genomes, including 113 newly assembled, we reconstructed the phylogenomic relationships, divergence times, biogeographic patterns, morphological evolution of basidiomata, and patterns of speciation/extinction in the Hymenochaetales. The phylogenomic relationships of 12 families within the Hymenochaetales suggested that 10 families can be accepted, and 2 families rejected. Molecular clock dating analyses suggested that the Hymenochaetales possibly started a rapid family-wide and genus-wide radiation during the early Cretaceous to late Jurassic and Cretaceous, respectively. Reconstruction of the ancestral state implied that Hymenochaetales probably originated from the temperate regions of Asia, with the basidiomata of the common ancestor likely being a corticioid species that rapidly transformed between the early Cretaceous and late Jurassic, coinciding with radiations at the family level. Furthermore, we detected a gradually increasing trend of speciation, extinction, and net diversification rates. We provided large-scale genomes of the Hymenochaetales and revealed evolutionary history patterns, which are key to understanding the evolution of fungi.
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spelling doaj-art-8e31a5b5c5034ad2830968fa5367e68c2025-08-20T02:25:41ZengTaylor & Francis GroupMycology2150-12032150-12112025-04-0116261763410.1080/21501203.2024.2391527Large-scale phylogenomic insights into the evolution of the HymenochaetalesHeng Zhao0Fang Wu1Sundy Maurice2Igor N. Pavlov3Konstantin V. Krutovsky4Hong-Gao Liu5Yuan Yuan6Yu-Cheng Dai7State Key Laboratory of Efficient Production of Forest Resources, School of Ecology and Nature Conservation, Beijing Forestry University, Beijing, ChinaState Key Laboratory of Efficient Production of Forest Resources, School of Ecology and Nature Conservation, Beijing Forestry University, Beijing, ChinaSection for Genetics and Evolutionary Biology (EVOGENE), Department of Biosciences, University of Oslo, Oslo, NorwayLaboratory of Mycology and Plant Pathology, V. N. Sukachev Institute of Forest, Siberian Branch, Russian Academy of Sciences, Krasnoyarsk, RussiaDepartment of Forest Genetics and Forest Tree Breeding, Georg-August University of Göttingen, Göttingen, GermanyYunnan Key Laboratory of Gastrodia and Fungi Symbiotic Biology, Zhaotong University, Zhaotong, ChinaState Key Laboratory of Efficient Production of Forest Resources, School of Ecology and Nature Conservation, Beijing Forestry University, Beijing, ChinaState Key Laboratory of Efficient Production of Forest Resources, School of Ecology and Nature Conservation, Beijing Forestry University, Beijing, ChinaThe Hymenochaetales is an order with most species as wood-inhabiting fungi that have high phylogenetic complexity and morphological diversity. Species in this order play important roles in forest ecosystems and include wood decomposers, pathogens, and those that form ectomycorrhizal associations. However, we have limited knowledge of the patterns of large-scale evolutionary history of the order. In this study, using 171 genomes, including 113 newly assembled, we reconstructed the phylogenomic relationships, divergence times, biogeographic patterns, morphological evolution of basidiomata, and patterns of speciation/extinction in the Hymenochaetales. The phylogenomic relationships of 12 families within the Hymenochaetales suggested that 10 families can be accepted, and 2 families rejected. Molecular clock dating analyses suggested that the Hymenochaetales possibly started a rapid family-wide and genus-wide radiation during the early Cretaceous to late Jurassic and Cretaceous, respectively. Reconstruction of the ancestral state implied that Hymenochaetales probably originated from the temperate regions of Asia, with the basidiomata of the common ancestor likely being a corticioid species that rapidly transformed between the early Cretaceous and late Jurassic, coinciding with radiations at the family level. Furthermore, we detected a gradually increasing trend of speciation, extinction, and net diversification rates. We provided large-scale genomes of the Hymenochaetales and revealed evolutionary history patterns, which are key to understanding the evolution of fungi.https://www.tandfonline.com/doi/10.1080/21501203.2024.2391527Ancestral statebiogeographic patternsHymenochaetaceaemolecular clock datingwhite-rot fungiwood-decay fungi
spellingShingle Heng Zhao
Fang Wu
Sundy Maurice
Igor N. Pavlov
Konstantin V. Krutovsky
Hong-Gao Liu
Yuan Yuan
Yu-Cheng Dai
Large-scale phylogenomic insights into the evolution of the Hymenochaetales
Mycology
Ancestral state
biogeographic patterns
Hymenochaetaceae
molecular clock dating
white-rot fungi
wood-decay fungi
title Large-scale phylogenomic insights into the evolution of the Hymenochaetales
title_full Large-scale phylogenomic insights into the evolution of the Hymenochaetales
title_fullStr Large-scale phylogenomic insights into the evolution of the Hymenochaetales
title_full_unstemmed Large-scale phylogenomic insights into the evolution of the Hymenochaetales
title_short Large-scale phylogenomic insights into the evolution of the Hymenochaetales
title_sort large scale phylogenomic insights into the evolution of the hymenochaetales
topic Ancestral state
biogeographic patterns
Hymenochaetaceae
molecular clock dating
white-rot fungi
wood-decay fungi
url https://www.tandfonline.com/doi/10.1080/21501203.2024.2391527
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