Structural characterization of glycinamide-RNase-transformylase T from Mycobacterium tuberculosis

Enzymes from the purine salvage pathway in Mycobacterium tuberculosis (Mtb) have been regarded as an attractive target for the development of anti-bacterial drugs. Although this pathway has not been extensively studied in Mtb, it has been identified as essential for growth and survival. Glycinamide-...

Full description

Saved in:
Bibliographic Details
Main Authors: Cong Chen, Zuliang Liu, Liguo Liu, Jianmin Wang, Qi Jin
Format: Article
Language:English
Published: Taylor & Francis Group 2020-01-01
Series:Emerging Microbes and Infections
Subjects:
Online Access:https://www.tandfonline.com/doi/10.1080/22221751.2019.1707716
Tags: Add Tag
No Tags, Be the first to tag this record!
_version_ 1850200418995404800
author Cong Chen
Zuliang Liu
Liguo Liu
Jianmin Wang
Qi Jin
author_facet Cong Chen
Zuliang Liu
Liguo Liu
Jianmin Wang
Qi Jin
author_sort Cong Chen
collection DOAJ
description Enzymes from the purine salvage pathway in Mycobacterium tuberculosis (Mtb) have been regarded as an attractive target for the development of anti-bacterial drugs. Although this pathway has not been extensively studied in Mtb, it has been identified as essential for growth and survival. Glycinamide-RNase-transformylase T (PurT) is found only in some specific bacteria including Mtb and utilizes ATP-dependent ligation to catalyze the formylation of 5′-phosphoribosyl-glycinamide (GAR) in the third reaction of the de novo purine salvage pathway. In the study, we determined the crystal structure of MtbPurT at a resolution of 2.79 Å. In contrast to Pyrococcus horikoshii OT3 PurT (phBCCPPurT), MtbPurT exhibits an “open” conformation, which results in a broader ATP-binding pocket and thus might facilitate the entry and exit of the cofactor. Additionally, active site superposition with E.coli PurT (EcPurT) showed that residues involved in the ATP-binding site in MtbPurT exhibited structural similarity but had notable difference in the GAR-binding site. The loop 383-389 in MtbPurT was much shorter and shifted 5.7 Å away from the phosphate of the GAR substrate. The different GAR-binding mode might result in a large conformational change in MtbPurT, and would provide a possible opportunity for anti-TB drug development.
format Article
id doaj-art-8bf8986a1e444ddfafec322f79ae36f0
institution OA Journals
issn 2222-1751
language English
publishDate 2020-01-01
publisher Taylor & Francis Group
record_format Article
series Emerging Microbes and Infections
spelling doaj-art-8bf8986a1e444ddfafec322f79ae36f02025-08-20T02:12:20ZengTaylor & Francis GroupEmerging Microbes and Infections2222-17512020-01-0191586610.1080/22221751.2019.1707716Structural characterization of glycinamide-RNase-transformylase T from Mycobacterium tuberculosisCong Chen0Zuliang Liu1Liguo Liu2Jianmin Wang3Qi Jin4NHC Key Laboratory of Systems Biology of Pathogens, Institute of Pathogen Biology, and Center for Tuberculosis Research, Chinese Academy of Medical Sciences and Peking Union Medical College, Beijing, P. R. People’s Republic of ChinaNHC Key Laboratory of Systems Biology of Pathogens, Institute of Pathogen Biology, and Center for Tuberculosis Research, Chinese Academy of Medical Sciences and Peking Union Medical College, Beijing, P. R. People’s Republic of ChinaNHC Key Laboratory of Systems Biology of Pathogens, Institute of Pathogen Biology, and Center for Tuberculosis Research, Chinese Academy of Medical Sciences and Peking Union Medical College, Beijing, P. R. People’s Republic of ChinaNHC Key Laboratory of Systems Biology of Pathogens, Institute of Pathogen Biology, and Center for Tuberculosis Research, Chinese Academy of Medical Sciences and Peking Union Medical College, Beijing, P. R. People’s Republic of ChinaNHC Key Laboratory of Systems Biology of Pathogens, Institute of Pathogen Biology, and Center for Tuberculosis Research, Chinese Academy of Medical Sciences and Peking Union Medical College, Beijing, P. R. People’s Republic of ChinaEnzymes from the purine salvage pathway in Mycobacterium tuberculosis (Mtb) have been regarded as an attractive target for the development of anti-bacterial drugs. Although this pathway has not been extensively studied in Mtb, it has been identified as essential for growth and survival. Glycinamide-RNase-transformylase T (PurT) is found only in some specific bacteria including Mtb and utilizes ATP-dependent ligation to catalyze the formylation of 5′-phosphoribosyl-glycinamide (GAR) in the third reaction of the de novo purine salvage pathway. In the study, we determined the crystal structure of MtbPurT at a resolution of 2.79 Å. In contrast to Pyrococcus horikoshii OT3 PurT (phBCCPPurT), MtbPurT exhibits an “open” conformation, which results in a broader ATP-binding pocket and thus might facilitate the entry and exit of the cofactor. Additionally, active site superposition with E.coli PurT (EcPurT) showed that residues involved in the ATP-binding site in MtbPurT exhibited structural similarity but had notable difference in the GAR-binding site. The loop 383-389 in MtbPurT was much shorter and shifted 5.7 Å away from the phosphate of the GAR substrate. The different GAR-binding mode might result in a large conformational change in MtbPurT, and would provide a possible opportunity for anti-TB drug development.https://www.tandfonline.com/doi/10.1080/22221751.2019.1707716Mycobacterium tuberculosisglycinamide-RNase-transformylase Tcrystal structurepurine salvage pathwayanti-TB drug development
spellingShingle Cong Chen
Zuliang Liu
Liguo Liu
Jianmin Wang
Qi Jin
Structural characterization of glycinamide-RNase-transformylase T from Mycobacterium tuberculosis
Emerging Microbes and Infections
Mycobacterium tuberculosis
glycinamide-RNase-transformylase T
crystal structure
purine salvage pathway
anti-TB drug development
title Structural characterization of glycinamide-RNase-transformylase T from Mycobacterium tuberculosis
title_full Structural characterization of glycinamide-RNase-transformylase T from Mycobacterium tuberculosis
title_fullStr Structural characterization of glycinamide-RNase-transformylase T from Mycobacterium tuberculosis
title_full_unstemmed Structural characterization of glycinamide-RNase-transformylase T from Mycobacterium tuberculosis
title_short Structural characterization of glycinamide-RNase-transformylase T from Mycobacterium tuberculosis
title_sort structural characterization of glycinamide rnase transformylase t from mycobacterium tuberculosis
topic Mycobacterium tuberculosis
glycinamide-RNase-transformylase T
crystal structure
purine salvage pathway
anti-TB drug development
url https://www.tandfonline.com/doi/10.1080/22221751.2019.1707716
work_keys_str_mv AT congchen structuralcharacterizationofglycinamidernasetransformylasetfrommycobacteriumtuberculosis
AT zuliangliu structuralcharacterizationofglycinamidernasetransformylasetfrommycobacteriumtuberculosis
AT liguoliu structuralcharacterizationofglycinamidernasetransformylasetfrommycobacteriumtuberculosis
AT jianminwang structuralcharacterizationofglycinamidernasetransformylasetfrommycobacteriumtuberculosis
AT qijin structuralcharacterizationofglycinamidernasetransformylasetfrommycobacteriumtuberculosis