Optineurin negatively regulates the induction of IFNbeta in response to RNA virus infection.

The innate immune response provides a critical defense against microbial infections, including viruses. These are recognised by pattern recognition receptors including Toll-like receptors (TLRs) and RIG-I like helicases (RLHs). Detection of virus triggers signalling cascades that induce transcriptio...

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Main Authors: Jamel Mankouri, Rennos Fragkoudis, Kathryn H Richards, Laura F Wetherill, Mark Harris, Alain Kohl, Richard M Elliott, Andrew Macdonald
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2010-02-01
Series:PLoS Pathogens
Online Access:https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1000778&type=printable
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author Jamel Mankouri
Rennos Fragkoudis
Kathryn H Richards
Laura F Wetherill
Mark Harris
Alain Kohl
Richard M Elliott
Andrew Macdonald
author_facet Jamel Mankouri
Rennos Fragkoudis
Kathryn H Richards
Laura F Wetherill
Mark Harris
Alain Kohl
Richard M Elliott
Andrew Macdonald
author_sort Jamel Mankouri
collection DOAJ
description The innate immune response provides a critical defense against microbial infections, including viruses. These are recognised by pattern recognition receptors including Toll-like receptors (TLRs) and RIG-I like helicases (RLHs). Detection of virus triggers signalling cascades that induce transcription of type I interferons including IFNbeta, which are pivotal for the initiation of an anti-viral state. Despite the essential role of IFNbeta in the anti-viral response, there is an incomplete understanding of the negative regulation of IFNbeta induction. Here we provide evidence that expression of the Nemo-related protein, optineurin (NRP/FIP2), has a role in the inhibition of virus-triggered IFNbeta induction. Over-expression of optineurin inhibited Sendai-virus (SeV) and dsRNA triggered induction of IFNbeta, whereas depletion of optineurin with siRNA promoted virus-induced IFNbeta production and decreased RNA virus replication. Immunoprecipitation and immunofluorescence studies identified optineurin in a protein complex containing the antiviral protein kinase TBK1 and the ubiquitin ligase TRAF3. Furthermore, mutagenesis studies determined that binding of ubiquitin was essential for both the correct sub-cellular localisation and the inhibitory function of optineurin. This work identifies optineurin as a critical regulator of antiviral signalling and potential target for future antiviral therapy.
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issn 1553-7366
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publishDate 2010-02-01
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spelling doaj-art-8b1cdff4e5f94af59bc041f47ac7542d2025-08-20T03:07:20ZengPublic Library of Science (PLoS)PLoS Pathogens1553-73661553-73742010-02-0162e100077810.1371/journal.ppat.1000778Optineurin negatively regulates the induction of IFNbeta in response to RNA virus infection.Jamel MankouriRennos FragkoudisKathryn H RichardsLaura F WetherillMark HarrisAlain KohlRichard M ElliottAndrew MacdonaldThe innate immune response provides a critical defense against microbial infections, including viruses. These are recognised by pattern recognition receptors including Toll-like receptors (TLRs) and RIG-I like helicases (RLHs). Detection of virus triggers signalling cascades that induce transcription of type I interferons including IFNbeta, which are pivotal for the initiation of an anti-viral state. Despite the essential role of IFNbeta in the anti-viral response, there is an incomplete understanding of the negative regulation of IFNbeta induction. Here we provide evidence that expression of the Nemo-related protein, optineurin (NRP/FIP2), has a role in the inhibition of virus-triggered IFNbeta induction. Over-expression of optineurin inhibited Sendai-virus (SeV) and dsRNA triggered induction of IFNbeta, whereas depletion of optineurin with siRNA promoted virus-induced IFNbeta production and decreased RNA virus replication. Immunoprecipitation and immunofluorescence studies identified optineurin in a protein complex containing the antiviral protein kinase TBK1 and the ubiquitin ligase TRAF3. Furthermore, mutagenesis studies determined that binding of ubiquitin was essential for both the correct sub-cellular localisation and the inhibitory function of optineurin. This work identifies optineurin as a critical regulator of antiviral signalling and potential target for future antiviral therapy.https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1000778&type=printable
spellingShingle Jamel Mankouri
Rennos Fragkoudis
Kathryn H Richards
Laura F Wetherill
Mark Harris
Alain Kohl
Richard M Elliott
Andrew Macdonald
Optineurin negatively regulates the induction of IFNbeta in response to RNA virus infection.
PLoS Pathogens
title Optineurin negatively regulates the induction of IFNbeta in response to RNA virus infection.
title_full Optineurin negatively regulates the induction of IFNbeta in response to RNA virus infection.
title_fullStr Optineurin negatively regulates the induction of IFNbeta in response to RNA virus infection.
title_full_unstemmed Optineurin negatively regulates the induction of IFNbeta in response to RNA virus infection.
title_short Optineurin negatively regulates the induction of IFNbeta in response to RNA virus infection.
title_sort optineurin negatively regulates the induction of ifnbeta in response to rna virus infection
url https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1000778&type=printable
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