Epigenetic Regulation of B Cell Memory Formation: A Poised Model for B Cell Epigenetic Reprograming

The formation of B cell immunological memory happens after the first encounter with a pathogen. At the germinal center (GC), B cells experience complex transcriptional and epigenetic transitions to differentiate into memory B cells (MBCs) and plasma cells (PCs). In particular, the differentiation of...

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Main Author: Carlos Valverde-Hernandez
Format: Article
Language:English
Published: Wiley 2025-01-01
Series:Journal of Immunology Research
Online Access:http://dx.doi.org/10.1155/jimr/9328523
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author Carlos Valverde-Hernandez
author_facet Carlos Valverde-Hernandez
author_sort Carlos Valverde-Hernandez
collection DOAJ
description The formation of B cell immunological memory happens after the first encounter with a pathogen. At the germinal center (GC), B cells experience complex transcriptional and epigenetic transitions to differentiate into memory B cells (MBCs) and plasma cells (PCs). In particular, the differentiation of GC B cells into MBCs has been poorly understood, and no clear conclusions on the signals and transcription factors leading to this cell fate have been identified. Recent discoveries in epigenetics and immune memory have elucidated the essential role of epigenetic regulators in establishing the memory B cell (MBC) fate. DNA methylation regulators, histone modifiers, noncoding RNAs (ncRNAs), and chromatin remodelers orchestrate a dynamic reprograming of the MBC phenotype. Positive and negative epigenetic regulators of the B cell program collaborate at each differentiation stage and allow for complex chromatin topology rearrangements and dynamic exposure to transcription and translation. Following MBC fate determination at the GC, the acquired epigenetic modifications induce a poised regulatory state where genes are epigenetically marked to remain transcriptionally inactive, but primed for rapid activation upon stimuli. Thus, a poised epigenetic control over gene expression governs MBC formation and a novel model of epigenetic reprograming is proposed. This model provides a novel perspective on how the B cell fate is determined in the GC and memory is formed, offering insights for improved vaccination and therapeutical approaches.
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spelling doaj-art-85400a9e3ee24a348c431e99b84dcb222025-08-20T02:45:49ZengWileyJournal of Immunology Research2314-71562025-01-01202510.1155/jimr/9328523Epigenetic Regulation of B Cell Memory Formation: A Poised Model for B Cell Epigenetic ReprogramingCarlos Valverde-Hernandez0Nuffield Department of Surgical SciencesThe formation of B cell immunological memory happens after the first encounter with a pathogen. At the germinal center (GC), B cells experience complex transcriptional and epigenetic transitions to differentiate into memory B cells (MBCs) and plasma cells (PCs). In particular, the differentiation of GC B cells into MBCs has been poorly understood, and no clear conclusions on the signals and transcription factors leading to this cell fate have been identified. Recent discoveries in epigenetics and immune memory have elucidated the essential role of epigenetic regulators in establishing the memory B cell (MBC) fate. DNA methylation regulators, histone modifiers, noncoding RNAs (ncRNAs), and chromatin remodelers orchestrate a dynamic reprograming of the MBC phenotype. Positive and negative epigenetic regulators of the B cell program collaborate at each differentiation stage and allow for complex chromatin topology rearrangements and dynamic exposure to transcription and translation. Following MBC fate determination at the GC, the acquired epigenetic modifications induce a poised regulatory state where genes are epigenetically marked to remain transcriptionally inactive, but primed for rapid activation upon stimuli. Thus, a poised epigenetic control over gene expression governs MBC formation and a novel model of epigenetic reprograming is proposed. This model provides a novel perspective on how the B cell fate is determined in the GC and memory is formed, offering insights for improved vaccination and therapeutical approaches.http://dx.doi.org/10.1155/jimr/9328523
spellingShingle Carlos Valverde-Hernandez
Epigenetic Regulation of B Cell Memory Formation: A Poised Model for B Cell Epigenetic Reprograming
Journal of Immunology Research
title Epigenetic Regulation of B Cell Memory Formation: A Poised Model for B Cell Epigenetic Reprograming
title_full Epigenetic Regulation of B Cell Memory Formation: A Poised Model for B Cell Epigenetic Reprograming
title_fullStr Epigenetic Regulation of B Cell Memory Formation: A Poised Model for B Cell Epigenetic Reprograming
title_full_unstemmed Epigenetic Regulation of B Cell Memory Formation: A Poised Model for B Cell Epigenetic Reprograming
title_short Epigenetic Regulation of B Cell Memory Formation: A Poised Model for B Cell Epigenetic Reprograming
title_sort epigenetic regulation of b cell memory formation a poised model for b cell epigenetic reprograming
url http://dx.doi.org/10.1155/jimr/9328523
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