A panoramic view of cotton resistance to Verticillium dahliae: From genetic architectures to precision genomic selection

Abstract Investigating the genetic regulatory mechanisms underlying complex traits forms the foundation for crop improvement. Verticillium wilt (VW), caused by Verticillium dahliae (V. dahliae), is one of the most devastating diseases affecting crop production worldwide. However, the genetic basis u...

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Main Authors: Xiaojun Zhang, Shiming Liu, Peng Wu, Wanying Xu, Dingyi Yang, Yuqing Ming, Shenghua Xiao, Weiran Wang, Jun Ma, Xinhui Nie, Zhan Gao, Junyuan Lv, Fei Wu, Zhaoguang Yang, Baoxin Zheng, Ping Du, Jiangmei Wang, Hao Ding, Jie Kong, Alifu Aierxi, Yu Yu, Wei Gao, Zhongxu Lin, Chunyuan You, Keith Lindsey, Nataša Štajner, Maojun Wang, Jiahe Wu, Shuangxia Jin, Xianlong Zhang, Longfu Zhu
Format: Article
Language:English
Published: Wiley 2025-06-01
Series:iMeta
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Online Access:https://doi.org/10.1002/imt2.70029
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author Xiaojun Zhang
Shiming Liu
Peng Wu
Wanying Xu
Dingyi Yang
Yuqing Ming
Shenghua Xiao
Weiran Wang
Jun Ma
Xinhui Nie
Zhan Gao
Junyuan Lv
Fei Wu
Zhaoguang Yang
Baoxin Zheng
Ping Du
Jiangmei Wang
Hao Ding
Jie Kong
Alifu Aierxi
Yu Yu
Wei Gao
Zhongxu Lin
Chunyuan You
Keith Lindsey
Nataša Štajner
Maojun Wang
Jiahe Wu
Shuangxia Jin
Xianlong Zhang
Longfu Zhu
author_facet Xiaojun Zhang
Shiming Liu
Peng Wu
Wanying Xu
Dingyi Yang
Yuqing Ming
Shenghua Xiao
Weiran Wang
Jun Ma
Xinhui Nie
Zhan Gao
Junyuan Lv
Fei Wu
Zhaoguang Yang
Baoxin Zheng
Ping Du
Jiangmei Wang
Hao Ding
Jie Kong
Alifu Aierxi
Yu Yu
Wei Gao
Zhongxu Lin
Chunyuan You
Keith Lindsey
Nataša Štajner
Maojun Wang
Jiahe Wu
Shuangxia Jin
Xianlong Zhang
Longfu Zhu
author_sort Xiaojun Zhang
collection DOAJ
description Abstract Investigating the genetic regulatory mechanisms underlying complex traits forms the foundation for crop improvement. Verticillium wilt (VW), caused by Verticillium dahliae (V. dahliae), is one of the most devastating diseases affecting crop production worldwide. However, the genetic basis underlying crop resistance to V. dahliae remains largely obscure, hindering progress in the genomic selection for VW resistance breeding. Here, we unraveled the genetic architectures and regulatory landscape of VW resistance in cotton by combining genome‐wide association studies (GWAS) and transcriptome‐wide association studies (TWAS) using 1152 transcriptomes derived from 290 cotton accessions. We identified 10 reliable quantitative trait loci (QTLs) associated with VW resistance across multiple environments. These QTLs showed a pyramiding resistance effect and exhibited promising efficacy in the genomic prediction of cotton's VW resistance supported by an F2:3 population. Moreover, trace analysis of these elite alleles revealed a notably increased utilization of Lsnp1, Lsnp4, Lsnp5, Lsnp8, and Lsnp9, which potentially contribute to the improvement of VW resistance in Chinese cotton breeding since the 1990s. We also identified remarkable gene modules and expression QTL (eQTL) hotspots related to the regulation of reactive oxygen species (ROS) homeostasis and immune response. Furthermore, 15 candidate causal genes were prioritized by TWAS. Knocking down eight genes with a negative effect significantly enhanced cotton resistance to V. dahliae. Among them, GhARM, encoding an armadillo (ARM)‐repeat protein, was verified to modulate cotton resistance to V. dahliae by regulating ROS homeostasis. Overall, this study updates the understanding of the genetic basis and regulatory mechanisms of cotton's VW resistance, providing valuable strategies for VW management through genomic selection in cotton breeding.
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spelling doaj-art-833c2ac127b14d4f9abbd89f0ab298862025-08-20T02:24:23ZengWileyiMeta2770-596X2025-06-0143n/an/a10.1002/imt2.70029A panoramic view of cotton resistance to Verticillium dahliae: From genetic architectures to precision genomic selectionXiaojun Zhang0Shiming Liu1Peng Wu2Wanying Xu3Dingyi Yang4Yuqing Ming5Shenghua Xiao6Weiran Wang7Jun Ma8Xinhui Nie9Zhan Gao10Junyuan Lv11Fei Wu12Zhaoguang Yang13Baoxin Zheng14Ping Du15Jiangmei Wang16Hao Ding17Jie Kong18Alifu Aierxi19Yu Yu20Wei Gao21Zhongxu Lin22Chunyuan You23Keith Lindsey24Nataša Štajner25Maojun Wang26Jiahe Wu27Shuangxia Jin28Xianlong Zhang29Longfu Zhu30National Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaInstitute of Economic Crops Xinjiang Academy of Agricultural Sciences Urumqi ChinaCollege of Agriculture Shihezi University Shihezi ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaInstitute of Economic Crops Xinjiang Academy of Agricultural Sciences Urumqi ChinaInstitute of Economic Crops Xinjiang Academy of Agricultural Sciences Urumqi ChinaCotton Research Institute Xinjiang Academy of Agriculture and Reclamation Science Shihezi ChinaState Key Laboratory of Cotton Biology Henan University Kaifeng ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaDepartment of Biosciences Durham University Durham UKBiotechnical Faculty University of Ljubljana Ljubljana SloveniaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaState Key Laboratory of Plant Genomics, Institute of Microbiology Chinese Academy of Sciences Beijing ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaNational Key Laboratory of Crop Genetic Improvement Huazhong Agricultural University Wuhan ChinaAbstract Investigating the genetic regulatory mechanisms underlying complex traits forms the foundation for crop improvement. Verticillium wilt (VW), caused by Verticillium dahliae (V. dahliae), is one of the most devastating diseases affecting crop production worldwide. However, the genetic basis underlying crop resistance to V. dahliae remains largely obscure, hindering progress in the genomic selection for VW resistance breeding. Here, we unraveled the genetic architectures and regulatory landscape of VW resistance in cotton by combining genome‐wide association studies (GWAS) and transcriptome‐wide association studies (TWAS) using 1152 transcriptomes derived from 290 cotton accessions. We identified 10 reliable quantitative trait loci (QTLs) associated with VW resistance across multiple environments. These QTLs showed a pyramiding resistance effect and exhibited promising efficacy in the genomic prediction of cotton's VW resistance supported by an F2:3 population. Moreover, trace analysis of these elite alleles revealed a notably increased utilization of Lsnp1, Lsnp4, Lsnp5, Lsnp8, and Lsnp9, which potentially contribute to the improvement of VW resistance in Chinese cotton breeding since the 1990s. We also identified remarkable gene modules and expression QTL (eQTL) hotspots related to the regulation of reactive oxygen species (ROS) homeostasis and immune response. Furthermore, 15 candidate causal genes were prioritized by TWAS. Knocking down eight genes with a negative effect significantly enhanced cotton resistance to V. dahliae. Among them, GhARM, encoding an armadillo (ARM)‐repeat protein, was verified to modulate cotton resistance to V. dahliae by regulating ROS homeostasis. Overall, this study updates the understanding of the genetic basis and regulatory mechanisms of cotton's VW resistance, providing valuable strategies for VW management through genomic selection in cotton breeding.https://doi.org/10.1002/imt2.70029armadillo‐repeat proteincottongenomic selectiongenome‐wide association studiesROS homeostasistranscriptome‐wide association studies
spellingShingle Xiaojun Zhang
Shiming Liu
Peng Wu
Wanying Xu
Dingyi Yang
Yuqing Ming
Shenghua Xiao
Weiran Wang
Jun Ma
Xinhui Nie
Zhan Gao
Junyuan Lv
Fei Wu
Zhaoguang Yang
Baoxin Zheng
Ping Du
Jiangmei Wang
Hao Ding
Jie Kong
Alifu Aierxi
Yu Yu
Wei Gao
Zhongxu Lin
Chunyuan You
Keith Lindsey
Nataša Štajner
Maojun Wang
Jiahe Wu
Shuangxia Jin
Xianlong Zhang
Longfu Zhu
A panoramic view of cotton resistance to Verticillium dahliae: From genetic architectures to precision genomic selection
iMeta
armadillo‐repeat protein
cotton
genomic selection
genome‐wide association studies
ROS homeostasis
transcriptome‐wide association studies
title A panoramic view of cotton resistance to Verticillium dahliae: From genetic architectures to precision genomic selection
title_full A panoramic view of cotton resistance to Verticillium dahliae: From genetic architectures to precision genomic selection
title_fullStr A panoramic view of cotton resistance to Verticillium dahliae: From genetic architectures to precision genomic selection
title_full_unstemmed A panoramic view of cotton resistance to Verticillium dahliae: From genetic architectures to precision genomic selection
title_short A panoramic view of cotton resistance to Verticillium dahliae: From genetic architectures to precision genomic selection
title_sort panoramic view of cotton resistance to verticillium dahliae from genetic architectures to precision genomic selection
topic armadillo‐repeat protein
cotton
genomic selection
genome‐wide association studies
ROS homeostasis
transcriptome‐wide association studies
url https://doi.org/10.1002/imt2.70029
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