CPSF6 defines a conserved capsid interface that modulates HIV-1 replication.

The HIV-1 genome enters cells inside a shell comprised of capsid (CA) protein. Variation in CA sequence alters HIV-1 infectivity and escape from host restriction factors. However, apart from the Cyclophilin A-binding loop, CA has no known interfaces with which to interact with cellular cofactors. He...

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Main Authors: Amanda J Price, Adam J Fletcher, Torsten Schaller, Tom Elliott, KyeongEun Lee, Vineet N KewalRamani, Jason W Chin, Greg J Towers, Leo C James
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2012-01-01
Series:PLoS Pathogens
Online Access:https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1002896&type=printable
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author Amanda J Price
Adam J Fletcher
Torsten Schaller
Tom Elliott
KyeongEun Lee
Vineet N KewalRamani
Jason W Chin
Greg J Towers
Leo C James
author_facet Amanda J Price
Adam J Fletcher
Torsten Schaller
Tom Elliott
KyeongEun Lee
Vineet N KewalRamani
Jason W Chin
Greg J Towers
Leo C James
author_sort Amanda J Price
collection DOAJ
description The HIV-1 genome enters cells inside a shell comprised of capsid (CA) protein. Variation in CA sequence alters HIV-1 infectivity and escape from host restriction factors. However, apart from the Cyclophilin A-binding loop, CA has no known interfaces with which to interact with cellular cofactors. Here we describe a novel protein-protein interface in the N-terminal domain of HIV-1 CA, determined by X-ray crystallography, which mediates both viral restriction and host cofactor dependence. The interface is highly conserved across lentiviruses and is accessible in the context of a hexameric lattice. Mutation of the interface prevents binding to and restriction by CPSF6-358, a truncated cytosolic form of the RNA processing factor, cleavage and polyadenylation specific factor 6 (CPSF6). Furthermore, mutations that prevent CPSF6 binding also relieve dependence on nuclear entry cofactors TNPO3 and RanBP2. These results suggest that the HIV-1 capsid mediates direct host cofactor interactions to facilitate viral infection.
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institution Kabale University
issn 1553-7366
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language English
publishDate 2012-01-01
publisher Public Library of Science (PLoS)
record_format Article
series PLoS Pathogens
spelling doaj-art-7ebe256ec2ca4132a1a8a19a34467a082025-08-20T03:26:47ZengPublic Library of Science (PLoS)PLoS Pathogens1553-73661553-73742012-01-0188e100289610.1371/journal.ppat.1002896CPSF6 defines a conserved capsid interface that modulates HIV-1 replication.Amanda J PriceAdam J FletcherTorsten SchallerTom ElliottKyeongEun LeeVineet N KewalRamaniJason W ChinGreg J TowersLeo C JamesThe HIV-1 genome enters cells inside a shell comprised of capsid (CA) protein. Variation in CA sequence alters HIV-1 infectivity and escape from host restriction factors. However, apart from the Cyclophilin A-binding loop, CA has no known interfaces with which to interact with cellular cofactors. Here we describe a novel protein-protein interface in the N-terminal domain of HIV-1 CA, determined by X-ray crystallography, which mediates both viral restriction and host cofactor dependence. The interface is highly conserved across lentiviruses and is accessible in the context of a hexameric lattice. Mutation of the interface prevents binding to and restriction by CPSF6-358, a truncated cytosolic form of the RNA processing factor, cleavage and polyadenylation specific factor 6 (CPSF6). Furthermore, mutations that prevent CPSF6 binding also relieve dependence on nuclear entry cofactors TNPO3 and RanBP2. These results suggest that the HIV-1 capsid mediates direct host cofactor interactions to facilitate viral infection.https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1002896&type=printable
spellingShingle Amanda J Price
Adam J Fletcher
Torsten Schaller
Tom Elliott
KyeongEun Lee
Vineet N KewalRamani
Jason W Chin
Greg J Towers
Leo C James
CPSF6 defines a conserved capsid interface that modulates HIV-1 replication.
PLoS Pathogens
title CPSF6 defines a conserved capsid interface that modulates HIV-1 replication.
title_full CPSF6 defines a conserved capsid interface that modulates HIV-1 replication.
title_fullStr CPSF6 defines a conserved capsid interface that modulates HIV-1 replication.
title_full_unstemmed CPSF6 defines a conserved capsid interface that modulates HIV-1 replication.
title_short CPSF6 defines a conserved capsid interface that modulates HIV-1 replication.
title_sort cpsf6 defines a conserved capsid interface that modulates hiv 1 replication
url https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1002896&type=printable
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