CCDC22 mutations that impair COMMD binding cause attenuated 3C/Ritscher-Schinzel syndrome

Abstract The CCC complex, composed of CCDC22, CCDC93, and ten proteins of the COMMD family, coordinates several critical steps required to recycle internalized plasma membrane proteins from endosomes to the cell surface. CCC interacts with Retriever, a trimeric cargo recognition complex comprising V...

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Main Authors: Amika Singla, Carolyn Rogers, Mary-Joe Touma, Yassin El-Najjar, Alison Colley, Daniel J. Boesch, Daniel D. Billadeau, Jozef Gecz, Baoyu Chen, Ezra Burstein
Format: Article
Language:English
Published: BMC 2025-05-01
Series:BMC Medical Genomics
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Online Access:https://doi.org/10.1186/s12920-025-02168-7
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author Amika Singla
Carolyn Rogers
Mary-Joe Touma
Yassin El-Najjar
Alison Colley
Daniel J. Boesch
Daniel D. Billadeau
Jozef Gecz
Baoyu Chen
Ezra Burstein
author_facet Amika Singla
Carolyn Rogers
Mary-Joe Touma
Yassin El-Najjar
Alison Colley
Daniel J. Boesch
Daniel D. Billadeau
Jozef Gecz
Baoyu Chen
Ezra Burstein
author_sort Amika Singla
collection DOAJ
description Abstract The CCC complex, composed of CCDC22, CCDC93, and ten proteins of the COMMD family, coordinates several critical steps required to recycle internalized plasma membrane proteins from endosomes to the cell surface. CCC interacts with Retriever, a trimeric cargo recognition complex comprising VPS35L, VPS26C, and VPS29, and works closely with the WASH complex, a crucial regulator of branched actin polymerization at endosomal membranes. Mutations in genes encoding subunits of these three complexes, CCDC22, VPS35L, and WASHC5, have been linked with a developmental syndrome known as 3 C (cranio-cerebello-cardiac) or Ritscher-Schinzel syndrome. Here, we report a new CCDC22 missense mutation, p.E208K, that results in attenuated 3 C syndrome, without cardiac or neuroanatomical abnormalities. We show that this mutation impairs CCC complex assembly by disrupting a conserved interaction surface required for CCDC22-COMMD4 binding. We also review previously described cases and identify that CCDC22 p.P172R has a similar attenuated phenotype and impairs complex assembly in a similar fashion as p.E208K. The characterization of these mutations adds to our understanding of the clinical and molecular spectrum of these disorders.
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spelling doaj-art-78d871e4d3cd472ea438e9098122c0872025-08-20T02:38:31ZengBMCBMC Medical Genomics1755-87942025-05-0118111110.1186/s12920-025-02168-7CCDC22 mutations that impair COMMD binding cause attenuated 3C/Ritscher-Schinzel syndromeAmika Singla0Carolyn Rogers1Mary-Joe Touma2Yassin El-Najjar3Alison Colley4Daniel J. Boesch5Daniel D. Billadeau6Jozef Gecz7Baoyu Chen8Ezra Burstein9Department of Internal Medicine, Division of Digestive and Liver Diseases, University of Texas Southwestern Medical CenterGenetics of Learning Disability Service, Hunter GeneticsDepartment of Internal Medicine, Division of Digestive and Liver Diseases, University of Texas Southwestern Medical CenterDepartment of Internal Medicine, Division of Digestive and Liver Diseases, University of Texas Southwestern Medical CenterGenetics of Learning Disability Service, Hunter GeneticsRoy J. Carver Department of Biochemistry, Biophysics & Molecular Biology, Iowa State UniversityDivision of Oncology Research, Mayo Clinic College of MedicineNeurogenetics Research Program, School of Pediatrics and Reproductive Health, University of AdelaideRoy J. Carver Department of Biochemistry, Biophysics & Molecular Biology, Iowa State UniversityDepartment of Internal Medicine, Division of Digestive and Liver Diseases, University of Texas Southwestern Medical CenterAbstract The CCC complex, composed of CCDC22, CCDC93, and ten proteins of the COMMD family, coordinates several critical steps required to recycle internalized plasma membrane proteins from endosomes to the cell surface. CCC interacts with Retriever, a trimeric cargo recognition complex comprising VPS35L, VPS26C, and VPS29, and works closely with the WASH complex, a crucial regulator of branched actin polymerization at endosomal membranes. Mutations in genes encoding subunits of these three complexes, CCDC22, VPS35L, and WASHC5, have been linked with a developmental syndrome known as 3 C (cranio-cerebello-cardiac) or Ritscher-Schinzel syndrome. Here, we report a new CCDC22 missense mutation, p.E208K, that results in attenuated 3 C syndrome, without cardiac or neuroanatomical abnormalities. We show that this mutation impairs CCC complex assembly by disrupting a conserved interaction surface required for CCDC22-COMMD4 binding. We also review previously described cases and identify that CCDC22 p.P172R has a similar attenuated phenotype and impairs complex assembly in a similar fashion as p.E208K. The characterization of these mutations adds to our understanding of the clinical and molecular spectrum of these disorders.https://doi.org/10.1186/s12920-025-02168-73 C syndromeCCDC22COMMD proteinRetrieverRitscher-Schinzel syndrome
spellingShingle Amika Singla
Carolyn Rogers
Mary-Joe Touma
Yassin El-Najjar
Alison Colley
Daniel J. Boesch
Daniel D. Billadeau
Jozef Gecz
Baoyu Chen
Ezra Burstein
CCDC22 mutations that impair COMMD binding cause attenuated 3C/Ritscher-Schinzel syndrome
BMC Medical Genomics
3 C syndrome
CCDC22
COMMD protein
Retriever
Ritscher-Schinzel syndrome
title CCDC22 mutations that impair COMMD binding cause attenuated 3C/Ritscher-Schinzel syndrome
title_full CCDC22 mutations that impair COMMD binding cause attenuated 3C/Ritscher-Schinzel syndrome
title_fullStr CCDC22 mutations that impair COMMD binding cause attenuated 3C/Ritscher-Schinzel syndrome
title_full_unstemmed CCDC22 mutations that impair COMMD binding cause attenuated 3C/Ritscher-Schinzel syndrome
title_short CCDC22 mutations that impair COMMD binding cause attenuated 3C/Ritscher-Schinzel syndrome
title_sort ccdc22 mutations that impair commd binding cause attenuated 3c ritscher schinzel syndrome
topic 3 C syndrome
CCDC22
COMMD protein
Retriever
Ritscher-Schinzel syndrome
url https://doi.org/10.1186/s12920-025-02168-7
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