Enhancer Extrachromosomal Circular DNA ANKRD28 Elicits Drug Resistance via POU2F2‐Mediated Transcriptional Network in Multiple Myeloma

Abstract Multiple myeloma (MM) remains an incurable disease primarily due to the emergence of drug resistance, and the underlying mechanisms remain unclear. Extrachromosomal circular DNAs (eccDNAs) are prevalent in cancer genomes of both coding and non‐coding regions. However, the role of non‐coding...

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Main Authors: Binzhen Chen, Jia Liu, Yaoxin Zhang, Changming Shi, Di Zhu, Guoqiang Zhang, Fei Xiao, Lu Zhong, Minyue Zhang, Lai Guan Ng, Honghui Huang, Tingting Lu, Jian Hou
Format: Article
Language:English
Published: Wiley 2025-06-01
Series:Advanced Science
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Online Access:https://doi.org/10.1002/advs.202415695
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author Binzhen Chen
Jia Liu
Yaoxin Zhang
Changming Shi
Di Zhu
Guoqiang Zhang
Fei Xiao
Lu Zhong
Minyue Zhang
Lai Guan Ng
Honghui Huang
Tingting Lu
Jian Hou
author_facet Binzhen Chen
Jia Liu
Yaoxin Zhang
Changming Shi
Di Zhu
Guoqiang Zhang
Fei Xiao
Lu Zhong
Minyue Zhang
Lai Guan Ng
Honghui Huang
Tingting Lu
Jian Hou
author_sort Binzhen Chen
collection DOAJ
description Abstract Multiple myeloma (MM) remains an incurable disease primarily due to the emergence of drug resistance, and the underlying mechanisms remain unclear. Extrachromosomal circular DNAs (eccDNAs) are prevalent in cancer genomes of both coding and non‐coding regions. However, the role of non‐coding eccDNA regions that serve as enhancers has been largely overlooked. Here, genome‐wide profiling of serum eccDNAs from donors and MM patients who responded well or poorly to bortezomib‐lenalidomide‐dexamethasone (VRd) therapy is characterized. A high copy number of eccDNA ANKRD28 (eccANKRD28) predicts poor therapy response and prognosis but enhanced transcriptional activity. Established VRd‐resistant MM cell lines exhibit a higher abundance of eccANKRD28, and CRISPR/Cas9‐mediated elevation of eccANKRD28 desensitizes bortezomib and lenalidomide treatment both in vitro and in vivo. Integrated multi‐omics analysis (H3K27ac ChIP‐seq, scRNA‐seq, scATAC‐seq, CUT&Tag, et al.) identifies eccANKRD28 as an active enhancer involved in drug resistance driven by the key transcription factor, POU class 2 homeobox 2 (POU2F2). POU2F2 interacts with sequence‐specific eccANKRD28 as well as RUNX1 and RUNX2 motifs to form the protein complex, which activates the promoter of oncogenes, including IRF4, JUNB, IKZF3, RUNX3, and BCL2. This study elucidates the potential transcriptional network of enhancer eccANKRD28 in MM drug resistance from a previously unrecognized epigenetic perspective.
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spelling doaj-art-782c19a5a7954fe396b3c8de291f32fd2025-08-20T02:02:58ZengWileyAdvanced Science2198-38442025-06-011221n/an/a10.1002/advs.202415695Enhancer Extrachromosomal Circular DNA ANKRD28 Elicits Drug Resistance via POU2F2‐Mediated Transcriptional Network in Multiple MyelomaBinzhen Chen0Jia Liu1Yaoxin Zhang2Changming Shi3Di Zhu4Guoqiang Zhang5Fei Xiao6Lu Zhong7Minyue Zhang8Lai Guan Ng9Honghui Huang10Tingting Lu11Jian Hou12Department of HematologyRenji HospitalShanghai Jiao Tong University School of MedicineShanghai 200127 ChinaDepartment of HematologyRenji HospitalShanghai Jiao Tong University School of MedicineShanghai 200127 ChinaNational Center for Gene Research National Key Laboratory of Plant Molecular Genetics CAS Center for Excellence in Molecular Plant Sciences Institute of Plant Physiology and Ecology Chinese Academy of Sciences Shanghai 200032 ChinaShanghai Immune Therapy Institute Renji Hospital Shanghai Jiao Tong University School of Medicine Shanghai 200127 ChinaDepartment of HematologyRenji HospitalShanghai Jiao Tong University School of MedicineShanghai 200127 ChinaDepartment of HematologyRenji HospitalShanghai Jiao Tong University School of MedicineShanghai 200127 ChinaDepartment of HematologyRenji HospitalShanghai Jiao Tong University School of MedicineShanghai 200127 ChinaDepartment of HematologyRenji HospitalShanghai Jiao Tong University School of MedicineShanghai 200127 ChinaDepartment of HematologyRenji HospitalShanghai Jiao Tong University School of MedicineShanghai 200127 ChinaShanghai Immune Therapy Institute Renji Hospital Shanghai Jiao Tong University School of Medicine Shanghai 200127 ChinaDepartment of HematologyRenji HospitalShanghai Jiao Tong University School of MedicineShanghai 200127 ChinaSheng Yushou Center of Cell Biology and Immunology Department of Genetics and Developmental Science School of Life Sciences and Biotechnology Shanghai Jiao Tong University Shanghai 200240 ChinaDepartment of HematologyRenji HospitalShanghai Jiao Tong University School of MedicineShanghai 200127 ChinaAbstract Multiple myeloma (MM) remains an incurable disease primarily due to the emergence of drug resistance, and the underlying mechanisms remain unclear. Extrachromosomal circular DNAs (eccDNAs) are prevalent in cancer genomes of both coding and non‐coding regions. However, the role of non‐coding eccDNA regions that serve as enhancers has been largely overlooked. Here, genome‐wide profiling of serum eccDNAs from donors and MM patients who responded well or poorly to bortezomib‐lenalidomide‐dexamethasone (VRd) therapy is characterized. A high copy number of eccDNA ANKRD28 (eccANKRD28) predicts poor therapy response and prognosis but enhanced transcriptional activity. Established VRd‐resistant MM cell lines exhibit a higher abundance of eccANKRD28, and CRISPR/Cas9‐mediated elevation of eccANKRD28 desensitizes bortezomib and lenalidomide treatment both in vitro and in vivo. Integrated multi‐omics analysis (H3K27ac ChIP‐seq, scRNA‐seq, scATAC‐seq, CUT&Tag, et al.) identifies eccANKRD28 as an active enhancer involved in drug resistance driven by the key transcription factor, POU class 2 homeobox 2 (POU2F2). POU2F2 interacts with sequence‐specific eccANKRD28 as well as RUNX1 and RUNX2 motifs to form the protein complex, which activates the promoter of oncogenes, including IRF4, JUNB, IKZF3, RUNX3, and BCL2. This study elucidates the potential transcriptional network of enhancer eccANKRD28 in MM drug resistance from a previously unrecognized epigenetic perspective.https://doi.org/10.1002/advs.202415695multiple myelomaextrachromosomal circular DNAdrug resistanceenhancertranscriptional regulation
spellingShingle Binzhen Chen
Jia Liu
Yaoxin Zhang
Changming Shi
Di Zhu
Guoqiang Zhang
Fei Xiao
Lu Zhong
Minyue Zhang
Lai Guan Ng
Honghui Huang
Tingting Lu
Jian Hou
Enhancer Extrachromosomal Circular DNA ANKRD28 Elicits Drug Resistance via POU2F2‐Mediated Transcriptional Network in Multiple Myeloma
Advanced Science
multiple myeloma
extrachromosomal circular DNA
drug resistance
enhancer
transcriptional regulation
title Enhancer Extrachromosomal Circular DNA ANKRD28 Elicits Drug Resistance via POU2F2‐Mediated Transcriptional Network in Multiple Myeloma
title_full Enhancer Extrachromosomal Circular DNA ANKRD28 Elicits Drug Resistance via POU2F2‐Mediated Transcriptional Network in Multiple Myeloma
title_fullStr Enhancer Extrachromosomal Circular DNA ANKRD28 Elicits Drug Resistance via POU2F2‐Mediated Transcriptional Network in Multiple Myeloma
title_full_unstemmed Enhancer Extrachromosomal Circular DNA ANKRD28 Elicits Drug Resistance via POU2F2‐Mediated Transcriptional Network in Multiple Myeloma
title_short Enhancer Extrachromosomal Circular DNA ANKRD28 Elicits Drug Resistance via POU2F2‐Mediated Transcriptional Network in Multiple Myeloma
title_sort enhancer extrachromosomal circular dna ankrd28 elicits drug resistance via pou2f2 mediated transcriptional network in multiple myeloma
topic multiple myeloma
extrachromosomal circular DNA
drug resistance
enhancer
transcriptional regulation
url https://doi.org/10.1002/advs.202415695
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