Integrated transcriptomic and metabolomic analysis of flavonoid biosynthesis in cigar tobacco leaves under variable nitrogen regimes

IntroductionFlavonoids are the most abundant secondary metabolites in plants and play important roles in plant growth, environmental adaptation, and human health. Tobacco serves as a vital cash crop and key source of flavonoids. Given tobacco's high nitrogen demand and the particular sensitivit...

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Main Authors: Kai Jia, Juanjuan Shi, Lingli Bai, Xueren Wang, Yuemin Wang, Xiangzhen Li, Wenqing Li, Chaoyuan Zheng
Format: Article
Language:English
Published: Frontiers Media S.A. 2025-06-01
Series:Frontiers in Plant Science
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Online Access:https://www.frontiersin.org/articles/10.3389/fpls.2025.1589215/full
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author Kai Jia
Juanjuan Shi
Lingli Bai
Xueren Wang
Yuemin Wang
Xiangzhen Li
Wenqing Li
Chaoyuan Zheng
author_facet Kai Jia
Juanjuan Shi
Lingli Bai
Xueren Wang
Yuemin Wang
Xiangzhen Li
Wenqing Li
Chaoyuan Zheng
author_sort Kai Jia
collection DOAJ
description IntroductionFlavonoids are the most abundant secondary metabolites in plants and play important roles in plant growth, environmental adaptation, and human health. Tobacco serves as a vital cash crop and key source of flavonoids. Given tobacco's high nitrogen demand and the particular sensitivity of flavonoid synthesis to nitrogen supply, understanding the regulatory mechanisms of flavonoid biosynthesis under varying nitrogen conditions is essential.Methods and resultsWe systematically investigated the flavonoid biosynthesis pathways in the cigar tobacco cultivar Haiyan 204 under different nitrogen application rates using integrated transcriptomic and metabolomic analyses. This study identified 694 differentially expressed metabolites (DEMs) and 6,114 differentially expressed genes (DEGs). Integrated analyses revealed significant enrichment in the flavonoid biosynthesis and phenylpropanoid biosynthesis pathways. A regulatory network of flavonoid biosynthesis in response to nitrogen was constructed. Weighted gene co-expression network analysis (WGCNA) identified CHS2 as a key gene strongly associated with flavonoid biosynthesis, alongside its potential regulatory transcription factors MYB7, MYB9, bHLH14, and MYB_related1. The reliability of transcriptome data was validated via qPCR.DiscussionThese results provide a scientific basis for elucidating the biosynthesis mechanisms of tobacco flavonoids and identifying key nitrogen-responsive genes. The constructed regulatory network and identified transcriptional regulators offer critical insights into nitrogen-modulated flavonoid synthesis in cigar tobacco.
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spelling doaj-art-76e7ea6297464f7dafa200633bcb06ae2025-08-20T03:30:14ZengFrontiers Media S.A.Frontiers in Plant Science1664-462X2025-06-011610.3389/fpls.2025.15892151589215Integrated transcriptomic and metabolomic analysis of flavonoid biosynthesis in cigar tobacco leaves under variable nitrogen regimesKai Jia0Juanjuan Shi1Lingli Bai2Xueren Wang3Yuemin Wang4Xiangzhen Li5Wenqing Li6Chaoyuan Zheng7College of Resources and Environment, Fujian Agriculture and Forestry University, Fuzhou, Fujian, ChinaCollege of Resources and Environment, Fujian Agriculture and Forestry University, Fuzhou, Fujian, ChinaCollege of Resources and Environment, Fujian Agriculture and Forestry University, Fuzhou, Fujian, ChinaSanming Tobacco Sciences Institute, Sanming Tobacco Company of Fujian Province, Sanming, Fujian, ChinaInstitute of Tobacco Sciences, Fujian Provincial Tobacco Monopoly Bureau, Fuzhou, Fujian, ChinaCollege of Resources and Environment, Fujian Agriculture and Forestry University, Fuzhou, Fujian, ChinaInstitute of Tobacco Sciences, Fujian Provincial Tobacco Monopoly Bureau, Fuzhou, Fujian, ChinaCollege of Resources and Environment, Fujian Agriculture and Forestry University, Fuzhou, Fujian, ChinaIntroductionFlavonoids are the most abundant secondary metabolites in plants and play important roles in plant growth, environmental adaptation, and human health. Tobacco serves as a vital cash crop and key source of flavonoids. Given tobacco's high nitrogen demand and the particular sensitivity of flavonoid synthesis to nitrogen supply, understanding the regulatory mechanisms of flavonoid biosynthesis under varying nitrogen conditions is essential.Methods and resultsWe systematically investigated the flavonoid biosynthesis pathways in the cigar tobacco cultivar Haiyan 204 under different nitrogen application rates using integrated transcriptomic and metabolomic analyses. This study identified 694 differentially expressed metabolites (DEMs) and 6,114 differentially expressed genes (DEGs). Integrated analyses revealed significant enrichment in the flavonoid biosynthesis and phenylpropanoid biosynthesis pathways. A regulatory network of flavonoid biosynthesis in response to nitrogen was constructed. Weighted gene co-expression network analysis (WGCNA) identified CHS2 as a key gene strongly associated with flavonoid biosynthesis, alongside its potential regulatory transcription factors MYB7, MYB9, bHLH14, and MYB_related1. The reliability of transcriptome data was validated via qPCR.DiscussionThese results provide a scientific basis for elucidating the biosynthesis mechanisms of tobacco flavonoids and identifying key nitrogen-responsive genes. The constructed regulatory network and identified transcriptional regulators offer critical insights into nitrogen-modulated flavonoid synthesis in cigar tobacco.https://www.frontiersin.org/articles/10.3389/fpls.2025.1589215/fullnitrogentranscriptomicsmetabolomicsflavonoidscigar tobacco
spellingShingle Kai Jia
Juanjuan Shi
Lingli Bai
Xueren Wang
Yuemin Wang
Xiangzhen Li
Wenqing Li
Chaoyuan Zheng
Integrated transcriptomic and metabolomic analysis of flavonoid biosynthesis in cigar tobacco leaves under variable nitrogen regimes
Frontiers in Plant Science
nitrogen
transcriptomics
metabolomics
flavonoids
cigar tobacco
title Integrated transcriptomic and metabolomic analysis of flavonoid biosynthesis in cigar tobacco leaves under variable nitrogen regimes
title_full Integrated transcriptomic and metabolomic analysis of flavonoid biosynthesis in cigar tobacco leaves under variable nitrogen regimes
title_fullStr Integrated transcriptomic and metabolomic analysis of flavonoid biosynthesis in cigar tobacco leaves under variable nitrogen regimes
title_full_unstemmed Integrated transcriptomic and metabolomic analysis of flavonoid biosynthesis in cigar tobacco leaves under variable nitrogen regimes
title_short Integrated transcriptomic and metabolomic analysis of flavonoid biosynthesis in cigar tobacco leaves under variable nitrogen regimes
title_sort integrated transcriptomic and metabolomic analysis of flavonoid biosynthesis in cigar tobacco leaves under variable nitrogen regimes
topic nitrogen
transcriptomics
metabolomics
flavonoids
cigar tobacco
url https://www.frontiersin.org/articles/10.3389/fpls.2025.1589215/full
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