Unravelling the Dynamic Physiological and Metabolome Responses of Wheat (<i>Triticum aestivum</i> L.) to Saline–Alkaline Stress at the Seedling Stage

Background/Objectives: Understanding metabolome adjustment under saline–alkaline conditions is crucial for enhancing crop tolerance capacity and ensuring food security. Although soil salinization impairs wheat seedlings’ growth, metabolome plasticity under saline–alkaline stress remains poorly under...

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Main Authors: Wei Ren, Li Chen
Format: Article
Language:English
Published: MDPI AG 2025-06-01
Series:Metabolites
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Online Access:https://www.mdpi.com/2218-1989/15/7/430
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author Wei Ren
Li Chen
author_facet Wei Ren
Li Chen
author_sort Wei Ren
collection DOAJ
description Background/Objectives: Understanding metabolome adjustment under saline–alkaline conditions is crucial for enhancing crop tolerance capacity and ensuring food security. Although soil salinization impairs wheat seedlings’ growth, metabolome plasticity under saline–alkaline stress remains poorly understood. Here, we delved into dynamic physiological and metabolome shifts in wheat seedlings grown on SAS (saline–alkaline soil) on the 7th and 15th days post-germination (DPG). Methods: A self-developed and cultivated high-generation salt–alkali wheat variety (011) was grown on SAS and control soil, followed by comparative physiological, biochemical, and metabolomics analyses of seedlings. Results: The seedlings’ saline–alkaline stress responses were developmentally regulated with reduced growth, increasing accumulation of proline and soluble sugars, and differential antioxidant response. LC-MS-based global metabolomics analysis revealed significant metabolite profile differences, with 367 and 485 differential metabolites identified on the 7th and 15th DPG, respectively, between control and treatment. Upregulation of saccharides, flavonoids, organic acids (citrate cycle-related), phenolic acids, amino acids and derivatives, phytohormones, and sphingolipid metabolism was essential for seedlings’ growth on SAS. The key induced metabolites in seedlings grown on SAS include saccharic acid, trehalose, sucrose, glucose, L-citramalic acid, phellodendroside, scutellarin, anthranilate-1-<i>O</i>-sophoroside, lavandulifolioside, N-methyl-L-glutamate, etc. Up-regulated phytohormones include abscisic acid (3.8-fold, 7th DPG and 3.18-fold, 15th DPG), jasmonic acid (1.93-fold, 15th DPG), and jasmonoyl isoleucine (2.03-fold, 15th DPG). Conclusions: Our findings highlight the importance of ABA and jasmonic acid in regulating salt–alkali tolerance in wheat seedlings. Moreover, this study depicts key pathways involved in salt–alkali tolerance in wheat seedlings and unveils key DMs, offering resources for boosting wheat production on SAS.
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spelling doaj-art-75dfa4e4693149eaac9eee632006e4312025-08-20T02:47:17ZengMDPI AGMetabolites2218-19892025-06-0115743010.3390/metabo15070430Unravelling the Dynamic Physiological and Metabolome Responses of Wheat (<i>Triticum aestivum</i> L.) to Saline–Alkaline Stress at the Seedling StageWei Ren0Li Chen1State Key Laboratory of Ecological Safety and Sustainable Development in Arid Lands, Xinjiang Institute of Ecology and Geography, Chinese Academy of Sciences, Urumqi 830011, ChinaState Key Laboratory of Ecological Safety and Sustainable Development in Arid Lands, Xinjiang Institute of Ecology and Geography, Chinese Academy of Sciences, Urumqi 830011, ChinaBackground/Objectives: Understanding metabolome adjustment under saline–alkaline conditions is crucial for enhancing crop tolerance capacity and ensuring food security. Although soil salinization impairs wheat seedlings’ growth, metabolome plasticity under saline–alkaline stress remains poorly understood. Here, we delved into dynamic physiological and metabolome shifts in wheat seedlings grown on SAS (saline–alkaline soil) on the 7th and 15th days post-germination (DPG). Methods: A self-developed and cultivated high-generation salt–alkali wheat variety (011) was grown on SAS and control soil, followed by comparative physiological, biochemical, and metabolomics analyses of seedlings. Results: The seedlings’ saline–alkaline stress responses were developmentally regulated with reduced growth, increasing accumulation of proline and soluble sugars, and differential antioxidant response. LC-MS-based global metabolomics analysis revealed significant metabolite profile differences, with 367 and 485 differential metabolites identified on the 7th and 15th DPG, respectively, between control and treatment. Upregulation of saccharides, flavonoids, organic acids (citrate cycle-related), phenolic acids, amino acids and derivatives, phytohormones, and sphingolipid metabolism was essential for seedlings’ growth on SAS. The key induced metabolites in seedlings grown on SAS include saccharic acid, trehalose, sucrose, glucose, L-citramalic acid, phellodendroside, scutellarin, anthranilate-1-<i>O</i>-sophoroside, lavandulifolioside, N-methyl-L-glutamate, etc. Up-regulated phytohormones include abscisic acid (3.8-fold, 7th DPG and 3.18-fold, 15th DPG), jasmonic acid (1.93-fold, 15th DPG), and jasmonoyl isoleucine (2.03-fold, 15th DPG). Conclusions: Our findings highlight the importance of ABA and jasmonic acid in regulating salt–alkali tolerance in wheat seedlings. Moreover, this study depicts key pathways involved in salt–alkali tolerance in wheat seedlings and unveils key DMs, offering resources for boosting wheat production on SAS.https://www.mdpi.com/2218-1989/15/7/430wheatsalt–alkali soilphytohormonedifferential metabolitemetabolic pathwaysLC-MS
spellingShingle Wei Ren
Li Chen
Unravelling the Dynamic Physiological and Metabolome Responses of Wheat (<i>Triticum aestivum</i> L.) to Saline–Alkaline Stress at the Seedling Stage
Metabolites
wheat
salt–alkali soil
phytohormone
differential metabolite
metabolic pathways
LC-MS
title Unravelling the Dynamic Physiological and Metabolome Responses of Wheat (<i>Triticum aestivum</i> L.) to Saline–Alkaline Stress at the Seedling Stage
title_full Unravelling the Dynamic Physiological and Metabolome Responses of Wheat (<i>Triticum aestivum</i> L.) to Saline–Alkaline Stress at the Seedling Stage
title_fullStr Unravelling the Dynamic Physiological and Metabolome Responses of Wheat (<i>Triticum aestivum</i> L.) to Saline–Alkaline Stress at the Seedling Stage
title_full_unstemmed Unravelling the Dynamic Physiological and Metabolome Responses of Wheat (<i>Triticum aestivum</i> L.) to Saline–Alkaline Stress at the Seedling Stage
title_short Unravelling the Dynamic Physiological and Metabolome Responses of Wheat (<i>Triticum aestivum</i> L.) to Saline–Alkaline Stress at the Seedling Stage
title_sort unravelling the dynamic physiological and metabolome responses of wheat i triticum aestivum i l to saline alkaline stress at the seedling stage
topic wheat
salt–alkali soil
phytohormone
differential metabolite
metabolic pathways
LC-MS
url https://www.mdpi.com/2218-1989/15/7/430
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AT lichen unravellingthedynamicphysiologicalandmetabolomeresponsesofwheatitriticumaestivumiltosalinealkalinestressattheseedlingstage