An enzyme activation network reveals extensive regulatory crosstalk between metabolic pathways

Abstract Enzyme activation by cellular metabolites plays a pivotal role in regulating metabolic processes. Nevertheless, our comprehension of such activation events on a global network scale remains incomplete. In this study, we conducted a comprehensive investigation into the optimization of cell-i...

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Main Authors: Sultana Mohammed Al Zubaidi, Muhammad Ibtisam Nasar, Richard A Notebaart, Markus Ralser, Mohammad Tauqeer Alam
Format: Article
Language:English
Published: Springer Nature 2025-05-01
Series:Molecular Systems Biology
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Online Access:https://doi.org/10.1038/s44320-025-00111-7
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author Sultana Mohammed Al Zubaidi
Muhammad Ibtisam Nasar
Richard A Notebaart
Markus Ralser
Mohammad Tauqeer Alam
author_facet Sultana Mohammed Al Zubaidi
Muhammad Ibtisam Nasar
Richard A Notebaart
Markus Ralser
Mohammad Tauqeer Alam
author_sort Sultana Mohammed Al Zubaidi
collection DOAJ
description Abstract Enzyme activation by cellular metabolites plays a pivotal role in regulating metabolic processes. Nevertheless, our comprehension of such activation events on a global network scale remains incomplete. In this study, we conducted a comprehensive investigation into the optimization of cell-intrinsic activation interactions using Saccharomyces cerevisiae metabolic network as the basis of the analysis. To achieve this, we integrated a genome-scale metabolic model with cross-species enzyme kinetic data sourced from the BRENDA database, and to use this model as a basis to estimate the distribution of enzyme activators throughout the cellular network. Our findings indicate that the vast majority of biochemical pathways encompass enzyme activators, frequently originating from disparate pathways, thus revealing extensive regulatory crosstalk between metabolic pathways. Notably, activators have short pathway lengths, indicating they are activated quickly upon nutrient shifts, and in most instances, these activators target key enzymatic reactions to facilitate downstream metabolic processes. Interestingly, highly activated enzymes are substantially enriched with non-essential enzymes compared to their essential counterparts. This observation suggests that cells employ enzyme activators to finely regulate secondary metabolic pathways that are only required under specific conditions. Conversely, the activator metabolites themselves are more likely to be essential components, and their activation levels surpass those of non-essential activators. In summary, our study unveils the widespread importance of enzymatic activators and suggests that feed-forward activation of conditional metabolic pathways through essential metabolites mediates metabolic plasticity.
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spelling doaj-art-714a6c9be6fd4e55908ce50e805402bf2025-08-20T04:01:43ZengSpringer NatureMolecular Systems Biology1744-42922025-05-0121787088810.1038/s44320-025-00111-7An enzyme activation network reveals extensive regulatory crosstalk between metabolic pathwaysSultana Mohammed Al Zubaidi0Muhammad Ibtisam Nasar1Richard A Notebaart2Markus Ralser3Mohammad Tauqeer Alam4Department of Biology, College of Science, United Arab Emirates UniversityDepartment of Biology, College of Science, United Arab Emirates UniversityFood Microbiology, Wageningen University and ResearchDepartment of Biochemistry, Charité – Universitätsmedizin BerlinDepartment of Biology, College of Science, United Arab Emirates UniversityAbstract Enzyme activation by cellular metabolites plays a pivotal role in regulating metabolic processes. Nevertheless, our comprehension of such activation events on a global network scale remains incomplete. In this study, we conducted a comprehensive investigation into the optimization of cell-intrinsic activation interactions using Saccharomyces cerevisiae metabolic network as the basis of the analysis. To achieve this, we integrated a genome-scale metabolic model with cross-species enzyme kinetic data sourced from the BRENDA database, and to use this model as a basis to estimate the distribution of enzyme activators throughout the cellular network. Our findings indicate that the vast majority of biochemical pathways encompass enzyme activators, frequently originating from disparate pathways, thus revealing extensive regulatory crosstalk between metabolic pathways. Notably, activators have short pathway lengths, indicating they are activated quickly upon nutrient shifts, and in most instances, these activators target key enzymatic reactions to facilitate downstream metabolic processes. Interestingly, highly activated enzymes are substantially enriched with non-essential enzymes compared to their essential counterparts. This observation suggests that cells employ enzyme activators to finely regulate secondary metabolic pathways that are only required under specific conditions. Conversely, the activator metabolites themselves are more likely to be essential components, and their activation levels surpass those of non-essential activators. In summary, our study unveils the widespread importance of enzymatic activators and suggests that feed-forward activation of conditional metabolic pathways through essential metabolites mediates metabolic plasticity.https://doi.org/10.1038/s44320-025-00111-7Intracellular Activation NetworkMetabolic RegulationRegulatory CrosstalkMetabolic NetworkActivator Compounds
spellingShingle Sultana Mohammed Al Zubaidi
Muhammad Ibtisam Nasar
Richard A Notebaart
Markus Ralser
Mohammad Tauqeer Alam
An enzyme activation network reveals extensive regulatory crosstalk between metabolic pathways
Molecular Systems Biology
Intracellular Activation Network
Metabolic Regulation
Regulatory Crosstalk
Metabolic Network
Activator Compounds
title An enzyme activation network reveals extensive regulatory crosstalk between metabolic pathways
title_full An enzyme activation network reveals extensive regulatory crosstalk between metabolic pathways
title_fullStr An enzyme activation network reveals extensive regulatory crosstalk between metabolic pathways
title_full_unstemmed An enzyme activation network reveals extensive regulatory crosstalk between metabolic pathways
title_short An enzyme activation network reveals extensive regulatory crosstalk between metabolic pathways
title_sort enzyme activation network reveals extensive regulatory crosstalk between metabolic pathways
topic Intracellular Activation Network
Metabolic Regulation
Regulatory Crosstalk
Metabolic Network
Activator Compounds
url https://doi.org/10.1038/s44320-025-00111-7
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