Convergent evolution links molybdenum insertase domains with organism-specific sequences

Abstract In all domains of life, the biosynthesis of the pterin-based Molybdenum cofactor (Moco) is crucial. Molybdenum (Mo) becomes biologically active by integrating into a unique pyranopterin scaffold, forming Moco. The final two steps of Moco biosynthesis are catalyzed by the two-domain enzyme M...

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Main Authors: Miriam Rabenow, Eduard Haar, Katharina Schmidt, Robert Hänsch, Ralf R. Mendel, Kevin D. Oliphant
Format: Article
Language:English
Published: Nature Portfolio 2024-10-01
Series:Communications Biology
Online Access:https://doi.org/10.1038/s42003-024-07073-w
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author Miriam Rabenow
Eduard Haar
Katharina Schmidt
Robert Hänsch
Ralf R. Mendel
Kevin D. Oliphant
author_facet Miriam Rabenow
Eduard Haar
Katharina Schmidt
Robert Hänsch
Ralf R. Mendel
Kevin D. Oliphant
author_sort Miriam Rabenow
collection DOAJ
description Abstract In all domains of life, the biosynthesis of the pterin-based Molybdenum cofactor (Moco) is crucial. Molybdenum (Mo) becomes biologically active by integrating into a unique pyranopterin scaffold, forming Moco. The final two steps of Moco biosynthesis are catalyzed by the two-domain enzyme Mo insertase, linked by gene fusion in higher organisms. Despite well-understood Moco biosynthesis, the evolutionary significance of Mo insertase fusion remains unclear. Here, we present findings from Neurospora crassa that shed light on the critical role of Mo insertase fusion in eukaryotes. Substituting the linkage region with sequences from other species resulted in Moco deficiency, and separate expression of domains, as seen in lower organisms, failed to rescue deficient strains. Stepwise truncation and structural modeling revealed a crucial 20-amino acid sequence within the linkage region essential for fungal growth. Our findings highlight the evolutionary importance of gene fusion and specific sequence composition in eukaryotic Mo insertases.
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issn 2399-3642
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spelling doaj-art-701bceec99d346f5bd0dd2bd5a7bd08f2025-08-20T02:17:53ZengNature PortfolioCommunications Biology2399-36422024-10-017111210.1038/s42003-024-07073-wConvergent evolution links molybdenum insertase domains with organism-specific sequencesMiriam Rabenow0Eduard Haar1Katharina Schmidt2Robert Hänsch3Ralf R. Mendel4Kevin D. Oliphant5Department of Plant Biology, Technische Universität BraunschweigDepartment of Plant Biology, Technische Universität BraunschweigDepartment of Plant Biology, Technische Universität BraunschweigDepartment of Plant Biology, Technische Universität BraunschweigDepartment of Plant Biology, Technische Universität BraunschweigDepartment of Plant Biology, Technische Universität BraunschweigAbstract In all domains of life, the biosynthesis of the pterin-based Molybdenum cofactor (Moco) is crucial. Molybdenum (Mo) becomes biologically active by integrating into a unique pyranopterin scaffold, forming Moco. The final two steps of Moco biosynthesis are catalyzed by the two-domain enzyme Mo insertase, linked by gene fusion in higher organisms. Despite well-understood Moco biosynthesis, the evolutionary significance of Mo insertase fusion remains unclear. Here, we present findings from Neurospora crassa that shed light on the critical role of Mo insertase fusion in eukaryotes. Substituting the linkage region with sequences from other species resulted in Moco deficiency, and separate expression of domains, as seen in lower organisms, failed to rescue deficient strains. Stepwise truncation and structural modeling revealed a crucial 20-amino acid sequence within the linkage region essential for fungal growth. Our findings highlight the evolutionary importance of gene fusion and specific sequence composition in eukaryotic Mo insertases.https://doi.org/10.1038/s42003-024-07073-w
spellingShingle Miriam Rabenow
Eduard Haar
Katharina Schmidt
Robert Hänsch
Ralf R. Mendel
Kevin D. Oliphant
Convergent evolution links molybdenum insertase domains with organism-specific sequences
Communications Biology
title Convergent evolution links molybdenum insertase domains with organism-specific sequences
title_full Convergent evolution links molybdenum insertase domains with organism-specific sequences
title_fullStr Convergent evolution links molybdenum insertase domains with organism-specific sequences
title_full_unstemmed Convergent evolution links molybdenum insertase domains with organism-specific sequences
title_short Convergent evolution links molybdenum insertase domains with organism-specific sequences
title_sort convergent evolution links molybdenum insertase domains with organism specific sequences
url https://doi.org/10.1038/s42003-024-07073-w
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