Evolution of the SARS-CoV-2 spike protein in utilizing host transmembrane serine proteases

Summary: SARS-CoV-2 entry into host cells depends on proteolytic activation of the spike protein by host proteases, a process shaped by spike mutations that influence viral specificity and infectivity. Using human airway epithelial models, this study investigated how different SARS-CoV-2 variants in...

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Main Authors: Aleksandra Milewska, Luis Fernando Cofas-Vargas, Adolfo B. Poma, Krzysztof Pyrć
Format: Article
Language:English
Published: Elsevier 2025-09-01
Series:iScience
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Online Access:http://www.sciencedirect.com/science/article/pii/S2589004225015792
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author Aleksandra Milewska
Luis Fernando Cofas-Vargas
Adolfo B. Poma
Krzysztof Pyrć
author_facet Aleksandra Milewska
Luis Fernando Cofas-Vargas
Adolfo B. Poma
Krzysztof Pyrć
author_sort Aleksandra Milewska
collection DOAJ
description Summary: SARS-CoV-2 entry into host cells depends on proteolytic activation of the spike protein by host proteases, a process shaped by spike mutations that influence viral specificity and infectivity. Using human airway epithelial models, this study investigated how different SARS-CoV-2 variants interact with host serine proteases. The Delta variant exhibited enhanced and stable binding to Hepsin through stronger ionic and hydrophobic interactions, promoting efficient spike activation and cell entry. In contrast, Omicron BA.1 showed weaker Hepsin binding and relied more on TMPRSS2 or cathepsins, depending on the cellular context. These findings reveal how variant-specific differences in protease usage are linked to spike protein mutations and cleavage site evolution. By illuminating the dynamic interplay between viral adaptation and host protease specificity, this work provides insights into mechanisms that influence viral transmission and immune evasion, with implications for developing targeted antiviral strategies and understanding the evolution of emerging SARS-CoV-2 variants.
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publishDate 2025-09-01
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spelling doaj-art-6cf07e3d60fc4f4b9941c28c1d9ff4042025-08-22T04:57:10ZengElsevieriScience2589-00422025-09-0128911331810.1016/j.isci.2025.113318Evolution of the SARS-CoV-2 spike protein in utilizing host transmembrane serine proteasesAleksandra Milewska0Luis Fernando Cofas-Vargas1Adolfo B. Poma2Krzysztof Pyrć3Virogenetics Laboratory of Virology, Malopolska Centre of Biotechnology, Jagiellonian University, Gronostajowa 7A, 30-387 Krakow, PolandInstitute of Fundamental Technological Research, Polish Academy of Sciences, Pawińskiego 5B, 02-106 Warsaw, PolandInstitute of Fundamental Technological Research, Polish Academy of Sciences, Pawińskiego 5B, 02-106 Warsaw, Poland; Corresponding authorVirogenetics Laboratory of Virology, Malopolska Centre of Biotechnology, Jagiellonian University, Gronostajowa 7A, 30-387 Krakow, Poland; Corresponding authorSummary: SARS-CoV-2 entry into host cells depends on proteolytic activation of the spike protein by host proteases, a process shaped by spike mutations that influence viral specificity and infectivity. Using human airway epithelial models, this study investigated how different SARS-CoV-2 variants interact with host serine proteases. The Delta variant exhibited enhanced and stable binding to Hepsin through stronger ionic and hydrophobic interactions, promoting efficient spike activation and cell entry. In contrast, Omicron BA.1 showed weaker Hepsin binding and relied more on TMPRSS2 or cathepsins, depending on the cellular context. These findings reveal how variant-specific differences in protease usage are linked to spike protein mutations and cleavage site evolution. By illuminating the dynamic interplay between viral adaptation and host protease specificity, this work provides insights into mechanisms that influence viral transmission and immune evasion, with implications for developing targeted antiviral strategies and understanding the evolution of emerging SARS-CoV-2 variants.http://www.sciencedirect.com/science/article/pii/S2589004225015792Molecular interactionCell biologyBioinformatics
spellingShingle Aleksandra Milewska
Luis Fernando Cofas-Vargas
Adolfo B. Poma
Krzysztof Pyrć
Evolution of the SARS-CoV-2 spike protein in utilizing host transmembrane serine proteases
iScience
Molecular interaction
Cell biology
Bioinformatics
title Evolution of the SARS-CoV-2 spike protein in utilizing host transmembrane serine proteases
title_full Evolution of the SARS-CoV-2 spike protein in utilizing host transmembrane serine proteases
title_fullStr Evolution of the SARS-CoV-2 spike protein in utilizing host transmembrane serine proteases
title_full_unstemmed Evolution of the SARS-CoV-2 spike protein in utilizing host transmembrane serine proteases
title_short Evolution of the SARS-CoV-2 spike protein in utilizing host transmembrane serine proteases
title_sort evolution of the sars cov 2 spike protein in utilizing host transmembrane serine proteases
topic Molecular interaction
Cell biology
Bioinformatics
url http://www.sciencedirect.com/science/article/pii/S2589004225015792
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