The transcription factor ZFP64 promotes activity-dependent synapse elimination during postnatal cerebellar development

Summary: Eliminating redundant synapses formed around birth is essential for shaping functionally mature neural circuits during postnatal development. Each Purkinje cell (PC) in the neonatal mouse cerebellum receives synaptic inputs from multiple climbing fibers (CFs). Only one CF is strengthened an...

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Main Authors: Jianling Zhang, Takaki Watanabe, Taisuke Miyazaki, Miwako Yamasaki, Kohtarou Konno, Yuto Okuno, Kyoko Matsuyama, Takayuki Noro, Masahiko Watanabe, Naofumi Uesaka, Masanobu Kano
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Language:English
Published: Elsevier 2025-06-01
Series:iScience
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Online Access:http://www.sciencedirect.com/science/article/pii/S2589004225010077
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author Jianling Zhang
Takaki Watanabe
Taisuke Miyazaki
Miwako Yamasaki
Kohtarou Konno
Yuto Okuno
Kyoko Matsuyama
Takayuki Noro
Masahiko Watanabe
Naofumi Uesaka
Masanobu Kano
author_facet Jianling Zhang
Takaki Watanabe
Taisuke Miyazaki
Miwako Yamasaki
Kohtarou Konno
Yuto Okuno
Kyoko Matsuyama
Takayuki Noro
Masahiko Watanabe
Naofumi Uesaka
Masanobu Kano
author_sort Jianling Zhang
collection DOAJ
description Summary: Eliminating redundant synapses formed around birth is essential for shaping functionally mature neural circuits during postnatal development. Each Purkinje cell (PC) in the neonatal mouse cerebellum receives synaptic inputs from multiple climbing fibers (CFs). Only one CF is strengthened and extends its innervation over PC dendrites, whereas the other CFs are eventually pruned during postnatal development. These events are believed to require proper gene expression, but the underlying mechanisms are not yet understood. Here, we report that the transcription factor ZFP64 in PCs mediates part of CF synapse elimination events presumably downstream of P/Q-type voltage-dependent Ca2+ channels (P/Q-VDCCs). PC-specific knockdown (KD) of ZFP64 during postnatal development delayed the elimination of redundant CF synapses and the dendritic extension of CF innervation. The KD of semaphorin 3A (Sema3A) in PCs partially restored the effects of ZFP64 or P/Q-VDCC KD. We propose that ZFP64 promotes developmental CF synapse elimination by regulating Sema3A expression.
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spelling doaj-art-6a19dcc7edf3403f925628fffcaeec712025-08-20T03:11:26ZengElsevieriScience2589-00422025-06-0128611274610.1016/j.isci.2025.112746The transcription factor ZFP64 promotes activity-dependent synapse elimination during postnatal cerebellar developmentJianling Zhang0Takaki Watanabe1Taisuke Miyazaki2Miwako Yamasaki3Kohtarou Konno4Yuto Okuno5Kyoko Matsuyama6Takayuki Noro7Masahiko Watanabe8Naofumi Uesaka9Masanobu Kano10Department of Neurophysiology, Graduate School of Medicine, The University of Tokyo, Tokyo 113-0033, Japan; International Research Center for Neurointelligence (WPI-IRCN), The University of Tokyo, Tokyo 113-0033, JapanDepartment of Neurophysiology, Graduate School of Medicine, The University of Tokyo, Tokyo 113-0033, Japan; International Research Center for Neurointelligence (WPI-IRCN), The University of Tokyo, Tokyo 113-0033, Japan; Advanced Comprehensive Research Organization (ACRO), Teikyo University, Tokyo 173-0003, Japan; Corresponding authorDepartment of Anatomy, Graduate School of Medicine, Hokkaido University, Sapporo 060-8638, JapanDepartment of Anatomy, Graduate School of Medicine, Hokkaido University, Sapporo 060-8638, JapanDepartment of Anatomy, Graduate School of Medicine, Hokkaido University, Sapporo 060-8638, JapanDepartment of Neurophysiology, Graduate School of Medicine, The University of Tokyo, Tokyo 113-0033, Japan; Advanced Comprehensive Research Organization (ACRO), Teikyo University, Tokyo 173-0003, JapanDepartment of Neurophysiology, Graduate School of Medicine, The University of Tokyo, Tokyo 113-0033, JapanDepartment of Neurophysiology, Graduate School of Medicine, The University of Tokyo, Tokyo 113-0033, JapanDepartment of Anatomy, Graduate School of Medicine, Hokkaido University, Sapporo 060-8638, JapanDepartment of Cognitive Neurobiology, Graduate School of Medical and Dental Sciences, Institute of Science Tokyo, Tokyo 113-8510, JapanDepartment of Neurophysiology, Graduate School of Medicine, The University of Tokyo, Tokyo 113-0033, Japan; International Research Center for Neurointelligence (WPI-IRCN), The University of Tokyo, Tokyo 113-0033, Japan; Advanced Comprehensive Research Organization (ACRO), Teikyo University, Tokyo 173-0003, Japan; Corresponding authorSummary: Eliminating redundant synapses formed around birth is essential for shaping functionally mature neural circuits during postnatal development. Each Purkinje cell (PC) in the neonatal mouse cerebellum receives synaptic inputs from multiple climbing fibers (CFs). Only one CF is strengthened and extends its innervation over PC dendrites, whereas the other CFs are eventually pruned during postnatal development. These events are believed to require proper gene expression, but the underlying mechanisms are not yet understood. Here, we report that the transcription factor ZFP64 in PCs mediates part of CF synapse elimination events presumably downstream of P/Q-type voltage-dependent Ca2+ channels (P/Q-VDCCs). PC-specific knockdown (KD) of ZFP64 during postnatal development delayed the elimination of redundant CF synapses and the dendritic extension of CF innervation. The KD of semaphorin 3A (Sema3A) in PCs partially restored the effects of ZFP64 or P/Q-VDCC KD. We propose that ZFP64 promotes developmental CF synapse elimination by regulating Sema3A expression.http://www.sciencedirect.com/science/article/pii/S2589004225010077Molecular physiologyNeuroscienceDevelopmental biology
spellingShingle Jianling Zhang
Takaki Watanabe
Taisuke Miyazaki
Miwako Yamasaki
Kohtarou Konno
Yuto Okuno
Kyoko Matsuyama
Takayuki Noro
Masahiko Watanabe
Naofumi Uesaka
Masanobu Kano
The transcription factor ZFP64 promotes activity-dependent synapse elimination during postnatal cerebellar development
iScience
Molecular physiology
Neuroscience
Developmental biology
title The transcription factor ZFP64 promotes activity-dependent synapse elimination during postnatal cerebellar development
title_full The transcription factor ZFP64 promotes activity-dependent synapse elimination during postnatal cerebellar development
title_fullStr The transcription factor ZFP64 promotes activity-dependent synapse elimination during postnatal cerebellar development
title_full_unstemmed The transcription factor ZFP64 promotes activity-dependent synapse elimination during postnatal cerebellar development
title_short The transcription factor ZFP64 promotes activity-dependent synapse elimination during postnatal cerebellar development
title_sort transcription factor zfp64 promotes activity dependent synapse elimination during postnatal cerebellar development
topic Molecular physiology
Neuroscience
Developmental biology
url http://www.sciencedirect.com/science/article/pii/S2589004225010077
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