TELP theory: Elucidating the major observations of Rieger et al. 2021 in mitochondria

The transmembrane-electrostatically localized protons (TELP) theory may represent a complementary development to Mitchell's chemiosmotic theory. The combination of the two together can now excellently explain the energetics in mitochondria. My calculated transmembrane-attractive force between a...

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Main Author: James Weifu Lee
Format: Article
Language:English
Published: KeAi Communications Co., Ltd. 2023-01-01
Series:Mitochondrial Communications
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Online Access:http://www.sciencedirect.com/science/article/pii/S2590279223000056
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author James Weifu Lee
author_facet James Weifu Lee
author_sort James Weifu Lee
collection DOAJ
description The transmembrane-electrostatically localized protons (TELP) theory may represent a complementary development to Mitchell's chemiosmotic theory. The combination of the two together can now excellently explain the energetics in mitochondria. My calculated transmembrane-attractive force between an excess proton and an excess hydroxide explains how TELP may stay within a 1-nm thin layer at the liquid-membrane interface. Consequently, any pH sensor (sEcGFP) located at least 2–3 nm away from the membrane surface will not be able to see TELP. This feature as predicted from the TELP model was observed exactly in the experiment of Rieger et al., 2021. In contrast to their belief “the Δp at ATP synthase is almost negligible under OXPHOS conditions”, I find, when TELP activity is included in the energy calculations, there is plenty of total protonic Gibbs free energy (ΔGT) well above the physiologically required value of −24.5 kJ mol−1 to drive ATP synthesis through FoF1-ATP synthase.
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spelling doaj-art-67e774a939a64b87b45e810a9280ed9f2025-08-20T03:56:14ZengKeAi Communications Co., Ltd.Mitochondrial Communications2590-27922023-01-011627210.1016/j.mitoco.2023.09.001TELP theory: Elucidating the major observations of Rieger et al. 2021 in mitochondriaJames Weifu Lee0Department of Chemistry and Biochemistry, Old Dominion University, Norfolk, VA, 23529, USAThe transmembrane-electrostatically localized protons (TELP) theory may represent a complementary development to Mitchell's chemiosmotic theory. The combination of the two together can now excellently explain the energetics in mitochondria. My calculated transmembrane-attractive force between an excess proton and an excess hydroxide explains how TELP may stay within a 1-nm thin layer at the liquid-membrane interface. Consequently, any pH sensor (sEcGFP) located at least 2–3 nm away from the membrane surface will not be able to see TELP. This feature as predicted from the TELP model was observed exactly in the experiment of Rieger et al., 2021. In contrast to their belief “the Δp at ATP synthase is almost negligible under OXPHOS conditions”, I find, when TELP activity is included in the energy calculations, there is plenty of total protonic Gibbs free energy (ΔGT) well above the physiologically required value of −24.5 kJ mol−1 to drive ATP synthesis through FoF1-ATP synthase.http://www.sciencedirect.com/science/article/pii/S2590279223000056Protonic capacitorWater as protonic conductorProton “hops and turns” mechanismExcess protonsTransmembrane-electrostatically localized protons (TELP)Local protonic motive force
spellingShingle James Weifu Lee
TELP theory: Elucidating the major observations of Rieger et al. 2021 in mitochondria
Mitochondrial Communications
Protonic capacitor
Water as protonic conductor
Proton “hops and turns” mechanism
Excess protons
Transmembrane-electrostatically localized protons (TELP)
Local protonic motive force
title TELP theory: Elucidating the major observations of Rieger et al. 2021 in mitochondria
title_full TELP theory: Elucidating the major observations of Rieger et al. 2021 in mitochondria
title_fullStr TELP theory: Elucidating the major observations of Rieger et al. 2021 in mitochondria
title_full_unstemmed TELP theory: Elucidating the major observations of Rieger et al. 2021 in mitochondria
title_short TELP theory: Elucidating the major observations of Rieger et al. 2021 in mitochondria
title_sort telp theory elucidating the major observations of rieger et al 2021 in mitochondria
topic Protonic capacitor
Water as protonic conductor
Proton “hops and turns” mechanism
Excess protons
Transmembrane-electrostatically localized protons (TELP)
Local protonic motive force
url http://www.sciencedirect.com/science/article/pii/S2590279223000056
work_keys_str_mv AT jamesweifulee telptheoryelucidatingthemajorobservationsofriegeretal2021inmitochondria