Sucrose synthase gene family in common bean during pod filling subjected to moisture restriction

In common bean (Phaseolus vulgaris L.), leaf photosynthesis is significantly reduced under drought conditions. Previous studies have shown that some drought-tolerant cultivars use the pod walls to compensate the decreased photosynthesis rate in leaves by acting as temporary reservoirs of carbohydrat...

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Main Authors: Norma Cecilia Morales-Elias, Eleazar Martínez-Barajas, Lilia Angélica Bernal-Gracida, Monserrat Vázquez-Sánchez, Iris Grisel Galván-Escobedo, Jose Salud Rodriguez-Zavala, Amelia López-Herrera, Cecilia Beatriz Peña-Valdivia, Antonio García-Esteva, Carlos Alberto Cruz-Cruz, María Teresa González-Arnao, José Cruz Jiménez-Galindo, Daniel Padilla-Chacón
Format: Article
Language:English
Published: Frontiers Media S.A. 2024-12-01
Series:Frontiers in Plant Science
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Online Access:https://www.frontiersin.org/articles/10.3389/fpls.2024.1462844/full
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author Norma Cecilia Morales-Elias
Eleazar Martínez-Barajas
Lilia Angélica Bernal-Gracida
Monserrat Vázquez-Sánchez
Iris Grisel Galván-Escobedo
Jose Salud Rodriguez-Zavala
Amelia López-Herrera
Cecilia Beatriz Peña-Valdivia
Antonio García-Esteva
Carlos Alberto Cruz-Cruz
María Teresa González-Arnao
José Cruz Jiménez-Galindo
Daniel Padilla-Chacón
author_facet Norma Cecilia Morales-Elias
Eleazar Martínez-Barajas
Lilia Angélica Bernal-Gracida
Monserrat Vázquez-Sánchez
Iris Grisel Galván-Escobedo
Jose Salud Rodriguez-Zavala
Amelia López-Herrera
Cecilia Beatriz Peña-Valdivia
Antonio García-Esteva
Carlos Alberto Cruz-Cruz
María Teresa González-Arnao
José Cruz Jiménez-Galindo
Daniel Padilla-Chacón
author_sort Norma Cecilia Morales-Elias
collection DOAJ
description In common bean (Phaseolus vulgaris L.), leaf photosynthesis is significantly reduced under drought conditions. Previous studies have shown that some drought-tolerant cultivars use the pod walls to compensate the decreased photosynthesis rate in leaves by acting as temporary reservoirs of carbohydrates to support seed filling. Here, we describe a comprehensive molecular characterization of sucrose synthase (SUS, EC 2.4.1.13) gene family through a genome-wide analysis and evaluated the effects of terminal drought on reproductive structures, specifically the pod walls. Seven PvSUS genes were located on six different chromosomes and had 8–16 intron–exon structures (8–16 exons). The PvSUS protein sequences revealed conserved catalytic domains, with molecular weights ranging from 90.5 kDa to 105.1 kDa and lengths from 799 to 929 amino acids. Phylogenetic analysis grouped these sequences into three main clusters with seven subgroups, indicating divergence from SUS sequences in other plant species. Using a docking sequence, we predicted three-dimensional (3-D) structures and evaluated the active sites. Bioinformatics analysis of promoter regions suggested that PvSUS genes may respond to light, hormone signaling, and stress stimuli. Greenhouse experiments were conducted using the cv. OTI, identified as having intermediate drought tolerance. Plants at the R8 growth stage were maintained with regular irrigation at 100% field capacity (FC) or with water restriction to maintain 50% of field capacity. Pods were harvested 5 days, 10 days, 15 days, and 20 days after anthesis. An increase in PvSUS activity under water restriction was associated with higher levels of fructose, while sucrose concentration also increased. qRT-PCR analysis revealed that PvSUS1, PvSUS3, and PvSUS4 were strongly expressed during seed development under water restriction. The fluorescent sucrose analog esculin indicated that transport across the plasma membrane might contribute to the increase in the pith cell diameter of pedicels. The results provide a systematic overview of the PvSUS gene family in P. vulgaris, offering a framework for further research and the potential functional application of PvSUS genes.
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spelling doaj-art-63cc21508ce34f9cbb184a55c2bd8f6b2025-08-20T01:58:03ZengFrontiers Media S.A.Frontiers in Plant Science1664-462X2024-12-011510.3389/fpls.2024.14628441462844Sucrose synthase gene family in common bean during pod filling subjected to moisture restrictionNorma Cecilia Morales-Elias0Eleazar Martínez-Barajas1Lilia Angélica Bernal-Gracida2Monserrat Vázquez-Sánchez3Iris Grisel Galván-Escobedo4Jose Salud Rodriguez-Zavala5Amelia López-Herrera6Cecilia Beatriz Peña-Valdivia7Antonio García-Esteva8Carlos Alberto Cruz-Cruz9María Teresa González-Arnao10José Cruz Jiménez-Galindo11Daniel Padilla-Chacón12Programa de Posgrado en Botánica, Colegio de Postgraduados Montecillo, Texcoco, MexicoFacultad de Química, Universidad Nacional Autónoma de México (UNAM), Mexico City, MexicoFacultad de Química, Universidad Nacional Autónoma de México (UNAM), Mexico City, MexicoPrograma de Posgrado en Botánica, Colegio de Postgraduados Montecillo, Texcoco, MexicoPrograma de Posgrado en Botánica, Colegio de Postgraduados Montecillo, Texcoco, MexicoDepartamento de Bioquímica, Instituto Nacional de Cardiología Ignacio Chávez, Mexico City, MexicoPosgrado en Recursos Genéticos y Productividad-Fisiología Vegetal, Colegio de Postgraduados Montecillo, Texcoco, MexicoPrograma de Posgrado en Botánica, Colegio de Postgraduados Montecillo, Texcoco, MexicoPrograma de Posgrado en Botánica, Colegio de Postgraduados Montecillo, Texcoco, MexicoFacultad de Ciencias Químicas, Universidad Veracruzana, Orizaba, Veracruz, MexicoFacultad de Ciencias Químicas, Universidad Veracruzana, Orizaba, Veracruz, MexicoSitio Experimental Sierra de Chihuahua-Instituto Nacional de Investigaciones Forestales, Agrícolas y Pecuarias (INIFAP), Chihuahua, MexicoConsejo Nacional de Humanidades, Ciencias y Tecnologías (CONAHCYT)-Facultad de Ciencias Químicas, Universidad Veracruzana, Orizaba, Veracruz, MexicoIn common bean (Phaseolus vulgaris L.), leaf photosynthesis is significantly reduced under drought conditions. Previous studies have shown that some drought-tolerant cultivars use the pod walls to compensate the decreased photosynthesis rate in leaves by acting as temporary reservoirs of carbohydrates to support seed filling. Here, we describe a comprehensive molecular characterization of sucrose synthase (SUS, EC 2.4.1.13) gene family through a genome-wide analysis and evaluated the effects of terminal drought on reproductive structures, specifically the pod walls. Seven PvSUS genes were located on six different chromosomes and had 8–16 intron–exon structures (8–16 exons). The PvSUS protein sequences revealed conserved catalytic domains, with molecular weights ranging from 90.5 kDa to 105.1 kDa and lengths from 799 to 929 amino acids. Phylogenetic analysis grouped these sequences into three main clusters with seven subgroups, indicating divergence from SUS sequences in other plant species. Using a docking sequence, we predicted three-dimensional (3-D) structures and evaluated the active sites. Bioinformatics analysis of promoter regions suggested that PvSUS genes may respond to light, hormone signaling, and stress stimuli. Greenhouse experiments were conducted using the cv. OTI, identified as having intermediate drought tolerance. Plants at the R8 growth stage were maintained with regular irrigation at 100% field capacity (FC) or with water restriction to maintain 50% of field capacity. Pods were harvested 5 days, 10 days, 15 days, and 20 days after anthesis. An increase in PvSUS activity under water restriction was associated with higher levels of fructose, while sucrose concentration also increased. qRT-PCR analysis revealed that PvSUS1, PvSUS3, and PvSUS4 were strongly expressed during seed development under water restriction. The fluorescent sucrose analog esculin indicated that transport across the plasma membrane might contribute to the increase in the pith cell diameter of pedicels. The results provide a systematic overview of the PvSUS gene family in P. vulgaris, offering a framework for further research and the potential functional application of PvSUS genes.https://www.frontiersin.org/articles/10.3389/fpls.2024.1462844/fullsucrosewater restrictionsucrose synthasePhaseolus vulgarispod wall
spellingShingle Norma Cecilia Morales-Elias
Eleazar Martínez-Barajas
Lilia Angélica Bernal-Gracida
Monserrat Vázquez-Sánchez
Iris Grisel Galván-Escobedo
Jose Salud Rodriguez-Zavala
Amelia López-Herrera
Cecilia Beatriz Peña-Valdivia
Antonio García-Esteva
Carlos Alberto Cruz-Cruz
María Teresa González-Arnao
José Cruz Jiménez-Galindo
Daniel Padilla-Chacón
Sucrose synthase gene family in common bean during pod filling subjected to moisture restriction
Frontiers in Plant Science
sucrose
water restriction
sucrose synthase
Phaseolus vulgaris
pod wall
title Sucrose synthase gene family in common bean during pod filling subjected to moisture restriction
title_full Sucrose synthase gene family in common bean during pod filling subjected to moisture restriction
title_fullStr Sucrose synthase gene family in common bean during pod filling subjected to moisture restriction
title_full_unstemmed Sucrose synthase gene family in common bean during pod filling subjected to moisture restriction
title_short Sucrose synthase gene family in common bean during pod filling subjected to moisture restriction
title_sort sucrose synthase gene family in common bean during pod filling subjected to moisture restriction
topic sucrose
water restriction
sucrose synthase
Phaseolus vulgaris
pod wall
url https://www.frontiersin.org/articles/10.3389/fpls.2024.1462844/full
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