Multimodal CRISPR screens uncover DDX39B as a global repressor of A-to-I RNA editing

Summary: Adenosine-to-inosine (A-to-I) RNA editing is a critical post-transcriptional modification that diversifies the transcriptome and influences various cellular processes, yet its regulatory mechanisms remain largely unknown. Here, we present two complementary CRISPR-based genetic screening pla...

Full description

Saved in:
Bibliographic Details
Main Authors: Tianzi Wei, Jiaxuan Li, Xiang Lei, Risheng Lin, Qingyan Wu, Zhenfeng Zhang, Shimin Shuai, Ruilin Tian
Format: Article
Language:English
Published: Elsevier 2025-07-01
Series:Cell Reports
Subjects:
Online Access:http://www.sciencedirect.com/science/article/pii/S2211124725007806
Tags: Add Tag
No Tags, Be the first to tag this record!
_version_ 1849703862980575232
author Tianzi Wei
Jiaxuan Li
Xiang Lei
Risheng Lin
Qingyan Wu
Zhenfeng Zhang
Shimin Shuai
Ruilin Tian
author_facet Tianzi Wei
Jiaxuan Li
Xiang Lei
Risheng Lin
Qingyan Wu
Zhenfeng Zhang
Shimin Shuai
Ruilin Tian
author_sort Tianzi Wei
collection DOAJ
description Summary: Adenosine-to-inosine (A-to-I) RNA editing is a critical post-transcriptional modification that diversifies the transcriptome and influences various cellular processes, yet its regulatory mechanisms remain largely unknown. Here, we present two complementary CRISPR-based genetic screening platforms: CREDITS (CRISPR-based RNA editing regulator screening), which enables genome-scale identification of editing regulators using an RNA recorder-based reporter system, and scCREDIT-seq (single-cell CRISPR-based RNA editing sequencing), which provides multiplexed single-cell characterization of transcriptome and editome changes for pooled perturbations. By screening 1,350 RNA-binding proteins, we identified a series of A-to-I editing regulators. Mechanistic investigation revealed DDX39B as a global repressor of A-to-I editing, which functions by preventing double-stranded RNA accumulation through its helicase activity. Targeting DDX39B significantly enhances the efficiency of RNA-editing-based tools, such as CellREADR (cell access through RNA sensing by endogenous ADAR) and LEAPER (leveraging endogenous ADAR for programmable editing of RNA), and disrupts hepatitis D virus (HDV) RNA editing homeostasis. These technological advances not only expand our understanding of RNA editing regulation but also provide powerful tools for exploring tissue-specific and context-dependent RNA modification mechanisms, with broad implications for therapeutic development.
format Article
id doaj-art-623e094d75bc48eb9dcbdaab350acabe
institution DOAJ
issn 2211-1247
language English
publishDate 2025-07-01
publisher Elsevier
record_format Article
series Cell Reports
spelling doaj-art-623e094d75bc48eb9dcbdaab350acabe2025-08-20T03:17:03ZengElsevierCell Reports2211-12472025-07-0144711600910.1016/j.celrep.2025.116009Multimodal CRISPR screens uncover DDX39B as a global repressor of A-to-I RNA editingTianzi Wei0Jiaxuan Li1Xiang Lei2Risheng Lin3Qingyan Wu4Zhenfeng Zhang5Shimin Shuai6Ruilin Tian7Department of Medical Neuroscience, School of Medicine, Southern University of Science and Technology, Shenzhen, Guangdong 518055, China; SUSTech Homeostatic Medicine Institute, School of Medicine, Southern University of Science and Technology, Shenzhen, Guangdong 518055, China; Key University Laboratory of Metabolism and Health of Guangdong, Southern University of Science and Technology, Shenzhen, Guangdong 518055, ChinaSUSTech Homeostatic Medicine Institute, School of Medicine, Southern University of Science and Technology, Shenzhen, Guangdong 518055, China; Department of Human Cell Biology and Genetics, School of Medicine, Southern University of Science and Technology, Shenzhen, Guangdong 518055, ChinaDepartment of Medical Neuroscience, School of Medicine, Southern University of Science and Technology, Shenzhen, Guangdong 518055, China; SUSTech Homeostatic Medicine Institute, School of Medicine, Southern University of Science and Technology, Shenzhen, Guangdong 518055, China; Key University Laboratory of Metabolism and Health of Guangdong, Southern University of Science and Technology, Shenzhen, Guangdong 518055, ChinaDepartment of Medical Neuroscience, School of Medicine, Southern University of Science and Technology, Shenzhen, Guangdong 518055, China; SUSTech Homeostatic Medicine Institute, School of Medicine, Southern University of Science and Technology, Shenzhen, Guangdong 518055, China; Key University Laboratory of Metabolism and Health of Guangdong, Southern University of Science and Technology, Shenzhen, Guangdong 518055, ChinaSchool of Public Health and Emergency Management, Southern University of Science and Technology, Shenzhen, Guangdong 518055, ChinaSchool of Public Health and Emergency Management, Southern University of Science and Technology, Shenzhen, Guangdong 518055, ChinaSUSTech Homeostatic Medicine Institute, School of Medicine, Southern University of Science and Technology, Shenzhen, Guangdong 518055, China; Department of Human Cell Biology and Genetics, School of Medicine, Southern University of Science and Technology, Shenzhen, Guangdong 518055, China; Corresponding authorDepartment of Medical Neuroscience, School of Medicine, Southern University of Science and Technology, Shenzhen, Guangdong 518055, China; SUSTech Homeostatic Medicine Institute, School of Medicine, Southern University of Science and Technology, Shenzhen, Guangdong 518055, China; Key University Laboratory of Metabolism and Health of Guangdong, Southern University of Science and Technology, Shenzhen, Guangdong 518055, China; Corresponding authorSummary: Adenosine-to-inosine (A-to-I) RNA editing is a critical post-transcriptional modification that diversifies the transcriptome and influences various cellular processes, yet its regulatory mechanisms remain largely unknown. Here, we present two complementary CRISPR-based genetic screening platforms: CREDITS (CRISPR-based RNA editing regulator screening), which enables genome-scale identification of editing regulators using an RNA recorder-based reporter system, and scCREDIT-seq (single-cell CRISPR-based RNA editing sequencing), which provides multiplexed single-cell characterization of transcriptome and editome changes for pooled perturbations. By screening 1,350 RNA-binding proteins, we identified a series of A-to-I editing regulators. Mechanistic investigation revealed DDX39B as a global repressor of A-to-I editing, which functions by preventing double-stranded RNA accumulation through its helicase activity. Targeting DDX39B significantly enhances the efficiency of RNA-editing-based tools, such as CellREADR (cell access through RNA sensing by endogenous ADAR) and LEAPER (leveraging endogenous ADAR for programmable editing of RNA), and disrupts hepatitis D virus (HDV) RNA editing homeostasis. These technological advances not only expand our understanding of RNA editing regulation but also provide powerful tools for exploring tissue-specific and context-dependent RNA modification mechanisms, with broad implications for therapeutic development.http://www.sciencedirect.com/science/article/pii/S2211124725007806CP: GenomicsCP: Molecular biology
spellingShingle Tianzi Wei
Jiaxuan Li
Xiang Lei
Risheng Lin
Qingyan Wu
Zhenfeng Zhang
Shimin Shuai
Ruilin Tian
Multimodal CRISPR screens uncover DDX39B as a global repressor of A-to-I RNA editing
Cell Reports
CP: Genomics
CP: Molecular biology
title Multimodal CRISPR screens uncover DDX39B as a global repressor of A-to-I RNA editing
title_full Multimodal CRISPR screens uncover DDX39B as a global repressor of A-to-I RNA editing
title_fullStr Multimodal CRISPR screens uncover DDX39B as a global repressor of A-to-I RNA editing
title_full_unstemmed Multimodal CRISPR screens uncover DDX39B as a global repressor of A-to-I RNA editing
title_short Multimodal CRISPR screens uncover DDX39B as a global repressor of A-to-I RNA editing
title_sort multimodal crispr screens uncover ddx39b as a global repressor of a to i rna editing
topic CP: Genomics
CP: Molecular biology
url http://www.sciencedirect.com/science/article/pii/S2211124725007806
work_keys_str_mv AT tianziwei multimodalcrisprscreensuncoverddx39basaglobalrepressorofatoirnaediting
AT jiaxuanli multimodalcrisprscreensuncoverddx39basaglobalrepressorofatoirnaediting
AT xianglei multimodalcrisprscreensuncoverddx39basaglobalrepressorofatoirnaediting
AT rishenglin multimodalcrisprscreensuncoverddx39basaglobalrepressorofatoirnaediting
AT qingyanwu multimodalcrisprscreensuncoverddx39basaglobalrepressorofatoirnaediting
AT zhenfengzhang multimodalcrisprscreensuncoverddx39basaglobalrepressorofatoirnaediting
AT shiminshuai multimodalcrisprscreensuncoverddx39basaglobalrepressorofatoirnaediting
AT ruilintian multimodalcrisprscreensuncoverddx39basaglobalrepressorofatoirnaediting