PRAG1 Condensation Drives Cell Contraction Under Stress

Peak1-related, kinase-activating pseudokinase 1 (PRAG1), a member of the pseudopodium-enriched atypical kinase (PEAK) family of pseudokinases, has been reported to play a role in regulating cell morphology. However, the molecular mechanism for this function remains elusive. In this study, we demonst...

Full description

Saved in:
Bibliographic Details
Main Authors: Peiwu Ye, Peiran Jiang, Luyu Ye, Min Liu, Qiuyuan Fang, Peilin Yu, Jianhong Luo, Huanxing Su, Wei Yang
Format: Article
Language:English
Published: MDPI AG 2025-03-01
Series:Biomolecules
Subjects:
Online Access:https://www.mdpi.com/2218-273X/15/3/379
Tags: Add Tag
No Tags, Be the first to tag this record!
_version_ 1850203969452769280
author Peiwu Ye
Peiran Jiang
Luyu Ye
Min Liu
Qiuyuan Fang
Peilin Yu
Jianhong Luo
Huanxing Su
Wei Yang
author_facet Peiwu Ye
Peiran Jiang
Luyu Ye
Min Liu
Qiuyuan Fang
Peilin Yu
Jianhong Luo
Huanxing Su
Wei Yang
author_sort Peiwu Ye
collection DOAJ
description Peak1-related, kinase-activating pseudokinase 1 (PRAG1), a member of the pseudopodium-enriched atypical kinase (PEAK) family of pseudokinases, has been reported to play a role in regulating cell morphology. However, the molecular mechanism for this function remains elusive. In this study, we demonstrate that PRAG1 forms dynamic condensates in cells mediated by its αN and αJ helices. Importantly, we found that PRAG1 condensates functioned in mediating cell contraction, while condensate-formation-deficient PRAG1 mutants lost this function. Remarkably, the formation of spherical PRAG1 condensates appears to be a common phenomenon in diverse stress models, as well as in dopaminergic (DA) neurons derived from a Parkinson’s disease patient. Our findings reveal a novel mechanism through which PRAG1 drives cell contraction and suggest a potential link between aberrant PRAG1 phase separation and stress-induced cell contraction. PRAG1 condensation drives cell contraction under stress.
format Article
id doaj-art-5ee33746b08641aeb6c134b8ae07c72c
institution OA Journals
issn 2218-273X
language English
publishDate 2025-03-01
publisher MDPI AG
record_format Article
series Biomolecules
spelling doaj-art-5ee33746b08641aeb6c134b8ae07c72c2025-08-20T02:11:22ZengMDPI AGBiomolecules2218-273X2025-03-0115337910.3390/biom15030379PRAG1 Condensation Drives Cell Contraction Under StressPeiwu Ye0Peiran Jiang1Luyu Ye2Min Liu3Qiuyuan Fang4Peilin Yu5Jianhong Luo6Huanxing Su7Wei Yang8Institute of Chinese Medical Sciences, University of Macau, Macau 999078, ChinaDepartment of Neurosurgery of the Fourth Affiliated Hospital, Zhejiang University School of Medicine, Hangzhou 310058, ChinaDepartment of Biophysics, Institute of Neuroscience, NHC and CAMS Key Laboratory of Medical Neurobiology, Zhejiang University School of Medicine, Hangzhou 310058, ChinaDepartment of Biophysics, Institute of Neuroscience, NHC and CAMS Key Laboratory of Medical Neurobiology, Zhejiang University School of Medicine, Hangzhou 310058, ChinaDepartment of Biophysics, Institute of Neuroscience, NHC and CAMS Key Laboratory of Medical Neurobiology, Zhejiang University School of Medicine, Hangzhou 310058, ChinaDepartment of Toxicology, School of Public Health, Zhejiang University School of Medicine, Hangzhou 310058, ChinaSchool of Brain Science and Brain Medicine, Zhejiang University, Hangzhou 310058, ChinaInstitute of Chinese Medical Sciences, University of Macau, Macau 999078, ChinaDepartment of Neurosurgery of the Fourth Affiliated Hospital, Zhejiang University School of Medicine, Hangzhou 310058, ChinaPeak1-related, kinase-activating pseudokinase 1 (PRAG1), a member of the pseudopodium-enriched atypical kinase (PEAK) family of pseudokinases, has been reported to play a role in regulating cell morphology. However, the molecular mechanism for this function remains elusive. In this study, we demonstrate that PRAG1 forms dynamic condensates in cells mediated by its αN and αJ helices. Importantly, we found that PRAG1 condensates functioned in mediating cell contraction, while condensate-formation-deficient PRAG1 mutants lost this function. Remarkably, the formation of spherical PRAG1 condensates appears to be a common phenomenon in diverse stress models, as well as in dopaminergic (DA) neurons derived from a Parkinson’s disease patient. Our findings reveal a novel mechanism through which PRAG1 drives cell contraction and suggest a potential link between aberrant PRAG1 phase separation and stress-induced cell contraction. PRAG1 condensation drives cell contraction under stress.https://www.mdpi.com/2218-273X/15/3/379PRAG1condensationphase separationcell contractionstress
spellingShingle Peiwu Ye
Peiran Jiang
Luyu Ye
Min Liu
Qiuyuan Fang
Peilin Yu
Jianhong Luo
Huanxing Su
Wei Yang
PRAG1 Condensation Drives Cell Contraction Under Stress
Biomolecules
PRAG1
condensation
phase separation
cell contraction
stress
title PRAG1 Condensation Drives Cell Contraction Under Stress
title_full PRAG1 Condensation Drives Cell Contraction Under Stress
title_fullStr PRAG1 Condensation Drives Cell Contraction Under Stress
title_full_unstemmed PRAG1 Condensation Drives Cell Contraction Under Stress
title_short PRAG1 Condensation Drives Cell Contraction Under Stress
title_sort prag1 condensation drives cell contraction under stress
topic PRAG1
condensation
phase separation
cell contraction
stress
url https://www.mdpi.com/2218-273X/15/3/379
work_keys_str_mv AT peiwuye prag1condensationdrivescellcontractionunderstress
AT peiranjiang prag1condensationdrivescellcontractionunderstress
AT luyuye prag1condensationdrivescellcontractionunderstress
AT minliu prag1condensationdrivescellcontractionunderstress
AT qiuyuanfang prag1condensationdrivescellcontractionunderstress
AT peilinyu prag1condensationdrivescellcontractionunderstress
AT jianhongluo prag1condensationdrivescellcontractionunderstress
AT huanxingsu prag1condensationdrivescellcontractionunderstress
AT weiyang prag1condensationdrivescellcontractionunderstress