Retrotransposon 3S18 forms self-protective aggregates and prolongs mid-oogenesis

Summary: Transposons are prevalent across nearly all species due to their capacity to mobilize in the host genome. However, their products may begin to affect the host before integration occurs. Here, we identified that the activation of transposons results in significantly smaller mid-stage oocytes...

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Main Authors: Dan Shen, Yaqian Xu, Qi Shi, Chongyang Li, Zhe Meng, Qiuju Wen, Chenhui Wang, Kun Dou
Format: Article
Language:English
Published: Elsevier 2025-07-01
Series:Cell Reports
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Online Access:http://www.sciencedirect.com/science/article/pii/S2211124725006850
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author Dan Shen
Yaqian Xu
Qi Shi
Chongyang Li
Zhe Meng
Qiuju Wen
Chenhui Wang
Kun Dou
author_facet Dan Shen
Yaqian Xu
Qi Shi
Chongyang Li
Zhe Meng
Qiuju Wen
Chenhui Wang
Kun Dou
author_sort Dan Shen
collection DOAJ
description Summary: Transposons are prevalent across nearly all species due to their capacity to mobilize in the host genome. However, their products may begin to affect the host before integration occurs. Here, we identified that the activation of transposons results in significantly smaller mid-stage oocytes and prolonged mid-oogenesis of Drosophila. Notably, one specific long terminal repeat (LTR) retrotransposon, 3S18, primarily contributes to this phenotype. We found that 3S18 mRNA and its integrase form micrometer-scaled ribonucleoprotein aggregates at cell-cell bridges during these stages. Interestingly, mutants that suppress the formation of these RNP aggregates substantially reduce 3S18 mRNA levels, suggesting that 3S18 aggregates serve functional importance in protecting the retrotransposon products. Live imaging reveals that the accumulation of 3S18 RNP aggregates obstructs host material transportation, resulting in prolonged mid-oogenesis. Finally, forcefully extending oogenesis significantly enhances 3S18 propagation. Our study highlights the unique characteristics of 3S18 and its impact on host development. It may shed light on studies of other parasitic elements, including viruses.
format Article
id doaj-art-5b6d46cfb64e48c3ae7bad7b34a136fd
institution Kabale University
issn 2211-1247
language English
publishDate 2025-07-01
publisher Elsevier
record_format Article
series Cell Reports
spelling doaj-art-5b6d46cfb64e48c3ae7bad7b34a136fd2025-08-20T03:24:08ZengElsevierCell Reports2211-12472025-07-0144711591410.1016/j.celrep.2025.115914Retrotransposon 3S18 forms self-protective aggregates and prolongs mid-oogenesisDan Shen0Yaqian Xu1Qi Shi2Chongyang Li3Zhe Meng4Qiuju Wen5Chenhui Wang6Kun Dou7School of Life Science and Technology, ShanghaiTech University, Shanghai 201210, ChinaSchool of Life Science and Technology, ShanghaiTech University, Shanghai 201210, ChinaSchool of Life Science and Technology, ShanghaiTech University, Shanghai 201210, ChinaSchool of Life Science and Technology, ShanghaiTech University, Shanghai 201210, ChinaSchool of Life Science and Technology, ShanghaiTech University, Shanghai 201210, ChinaSchool of Life Science and Technology, ShanghaiTech University, Shanghai 201210, ChinaSchool of Life Science and Technology, ShanghaiTech University, Shanghai 201210, ChinaSchool of Life Science and Technology, ShanghaiTech University, Shanghai 201210, China; Corresponding authorSummary: Transposons are prevalent across nearly all species due to their capacity to mobilize in the host genome. However, their products may begin to affect the host before integration occurs. Here, we identified that the activation of transposons results in significantly smaller mid-stage oocytes and prolonged mid-oogenesis of Drosophila. Notably, one specific long terminal repeat (LTR) retrotransposon, 3S18, primarily contributes to this phenotype. We found that 3S18 mRNA and its integrase form micrometer-scaled ribonucleoprotein aggregates at cell-cell bridges during these stages. Interestingly, mutants that suppress the formation of these RNP aggregates substantially reduce 3S18 mRNA levels, suggesting that 3S18 aggregates serve functional importance in protecting the retrotransposon products. Live imaging reveals that the accumulation of 3S18 RNP aggregates obstructs host material transportation, resulting in prolonged mid-oogenesis. Finally, forcefully extending oogenesis significantly enhances 3S18 propagation. Our study highlights the unique characteristics of 3S18 and its impact on host development. It may shed light on studies of other parasitic elements, including viruses.http://www.sciencedirect.com/science/article/pii/S2211124725006850CP: Molecular biologyCP: Developmental biology
spellingShingle Dan Shen
Yaqian Xu
Qi Shi
Chongyang Li
Zhe Meng
Qiuju Wen
Chenhui Wang
Kun Dou
Retrotransposon 3S18 forms self-protective aggregates and prolongs mid-oogenesis
Cell Reports
CP: Molecular biology
CP: Developmental biology
title Retrotransposon 3S18 forms self-protective aggregates and prolongs mid-oogenesis
title_full Retrotransposon 3S18 forms self-protective aggregates and prolongs mid-oogenesis
title_fullStr Retrotransposon 3S18 forms self-protective aggregates and prolongs mid-oogenesis
title_full_unstemmed Retrotransposon 3S18 forms self-protective aggregates and prolongs mid-oogenesis
title_short Retrotransposon 3S18 forms self-protective aggregates and prolongs mid-oogenesis
title_sort retrotransposon 3s18 forms self protective aggregates and prolongs mid oogenesis
topic CP: Molecular biology
CP: Developmental biology
url http://www.sciencedirect.com/science/article/pii/S2211124725006850
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AT chongyangli retrotransposon3s18formsselfprotectiveaggregatesandprolongsmidoogenesis
AT zhemeng retrotransposon3s18formsselfprotectiveaggregatesandprolongsmidoogenesis
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