Kinetic Approaches to Understanding the Mechanisms of Fidelity of the Herpes Simplex Virus Type 1 DNA Polymerase

We discuss how the results of presteady-state and steady-state kinetic analysis of the polymerizing and excision activities of herpes simplex virus type 1 (HSV-1) DNA polymerase have led to a better understanding of the mechanisms controlling fidelity of this important model replication polymerase....

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Main Authors: Yali Zhu, Jason Stroud, Liping Song, Deborah S. Parris
Format: Article
Language:English
Published: Wiley 2010-01-01
Series:Journal of Nucleic Acids
Online Access:http://dx.doi.org/10.4061/2010/631595
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author Yali Zhu
Jason Stroud
Liping Song
Deborah S. Parris
author_facet Yali Zhu
Jason Stroud
Liping Song
Deborah S. Parris
author_sort Yali Zhu
collection DOAJ
description We discuss how the results of presteady-state and steady-state kinetic analysis of the polymerizing and excision activities of herpes simplex virus type 1 (HSV-1) DNA polymerase have led to a better understanding of the mechanisms controlling fidelity of this important model replication polymerase. Despite a poorer misincorporation frequency compared to other replicative polymerases with intrinsic 3′ to 5′ exonuclease (exo) activity, HSV-1 DNA replication fidelity is enhanced by a high kinetic barrier to extending a primer/template containing a mismatch or abasic lesion and by the dynamic ability of the polymerase to switch the primer terminus between the exo and polymerizing active sites. The HSV-1 polymerase with a catalytically inactivated exo activity possesses reduced rates of primer switching and fails to support productive replication, suggesting a novel means to target polymerase for replication inhibition.
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spelling doaj-art-5622fe4e172544be9abde5dde31b0ce92025-08-20T02:04:58ZengWileyJournal of Nucleic Acids2090-021X2010-01-01201010.4061/2010/631595631595Kinetic Approaches to Understanding the Mechanisms of Fidelity of the Herpes Simplex Virus Type 1 DNA PolymeraseYali Zhu0Jason Stroud1Liping Song2Deborah S. Parris3Department of Molecular Virology, Immunology, and Medical Genetics, The Ohio State University, 2198 Graves Hall, 333 West Tenth Avenue, Columbus, OH 43210, USADepartment of Molecular Genetics, The Ohio State University, 2198 Graves Hall, 333 West Tenth Avenue, Columbus, OH 43210, USADepartment of Molecular Virology, Immunology, and Medical Genetics, The Ohio State University, 2198 Graves Hall, 333 West Tenth Avenue, Columbus, OH 43210, USADepartment of Molecular Virology, Immunology, and Medical Genetics, The Ohio State University, 2198 Graves Hall, 333 West Tenth Avenue, Columbus, OH 43210, USAWe discuss how the results of presteady-state and steady-state kinetic analysis of the polymerizing and excision activities of herpes simplex virus type 1 (HSV-1) DNA polymerase have led to a better understanding of the mechanisms controlling fidelity of this important model replication polymerase. Despite a poorer misincorporation frequency compared to other replicative polymerases with intrinsic 3′ to 5′ exonuclease (exo) activity, HSV-1 DNA replication fidelity is enhanced by a high kinetic barrier to extending a primer/template containing a mismatch or abasic lesion and by the dynamic ability of the polymerase to switch the primer terminus between the exo and polymerizing active sites. The HSV-1 polymerase with a catalytically inactivated exo activity possesses reduced rates of primer switching and fails to support productive replication, suggesting a novel means to target polymerase for replication inhibition.http://dx.doi.org/10.4061/2010/631595
spellingShingle Yali Zhu
Jason Stroud
Liping Song
Deborah S. Parris
Kinetic Approaches to Understanding the Mechanisms of Fidelity of the Herpes Simplex Virus Type 1 DNA Polymerase
Journal of Nucleic Acids
title Kinetic Approaches to Understanding the Mechanisms of Fidelity of the Herpes Simplex Virus Type 1 DNA Polymerase
title_full Kinetic Approaches to Understanding the Mechanisms of Fidelity of the Herpes Simplex Virus Type 1 DNA Polymerase
title_fullStr Kinetic Approaches to Understanding the Mechanisms of Fidelity of the Herpes Simplex Virus Type 1 DNA Polymerase
title_full_unstemmed Kinetic Approaches to Understanding the Mechanisms of Fidelity of the Herpes Simplex Virus Type 1 DNA Polymerase
title_short Kinetic Approaches to Understanding the Mechanisms of Fidelity of the Herpes Simplex Virus Type 1 DNA Polymerase
title_sort kinetic approaches to understanding the mechanisms of fidelity of the herpes simplex virus type 1 dna polymerase
url http://dx.doi.org/10.4061/2010/631595
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