Membrane lipid composition modulates the organization of VDAC1, a mitochondrial gatekeeper

Abstract VDACs, the most abundant proteins in the outer mitochondrial membrane (MOM), are crucial for mitochondrial physiology. VDAC regulate metabolite and ion exchange, modulate calcium homeostasis, and play roles in numerous cellular events such as apoptosis, mitochondrial DNA (mtDNA) release, an...

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Main Authors: Elodie Lafargue, Jean-Pierre Duneau, Nicolas Buzhinsky, Pamela Ornelas, Alexandre Ortega, Varun Ravishankar, James Sturgis, Ignacio Casuso, Lucie Bergdoll
Format: Article
Language:English
Published: Nature Portfolio 2025-06-01
Series:Communications Biology
Online Access:https://doi.org/10.1038/s42003-025-08311-5
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author Elodie Lafargue
Jean-Pierre Duneau
Nicolas Buzhinsky
Pamela Ornelas
Alexandre Ortega
Varun Ravishankar
James Sturgis
Ignacio Casuso
Lucie Bergdoll
author_facet Elodie Lafargue
Jean-Pierre Duneau
Nicolas Buzhinsky
Pamela Ornelas
Alexandre Ortega
Varun Ravishankar
James Sturgis
Ignacio Casuso
Lucie Bergdoll
author_sort Elodie Lafargue
collection DOAJ
description Abstract VDACs, the most abundant proteins in the outer mitochondrial membrane (MOM), are crucial for mitochondrial physiology. VDAC regulate metabolite and ion exchange, modulate calcium homeostasis, and play roles in numerous cellular events such as apoptosis, mitochondrial DNA (mtDNA) release, and different diseases. Mitochondrial function is closely tied to VDAC oligomerization, influencing key processes like mtDNA release and apoptosis, but the molecular drivers of this oligomerization remain unclear. In this study, we investigate the effects of three major MOM lipids on VDAC assemblies using atomic force microscopy and molecular dynamics simulations. Our results show that phosphatidylethanolamine and cholesterol regulate VDAC assembly, with the formation of stable lipid–protein organization of various size and compaction. Deviations from physiological lipid content disrupted native-like VDAC assemblies, highlighting the importance of lipid environment in VDAC organization. These findings underscore how lipid heterogeneity and changes in membranes influence VDAC function.
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publisher Nature Portfolio
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series Communications Biology
spelling doaj-art-4ec460e5e71a4da2a83fb95f399e01212025-08-20T02:37:35ZengNature PortfolioCommunications Biology2399-36422025-06-018111210.1038/s42003-025-08311-5Membrane lipid composition modulates the organization of VDAC1, a mitochondrial gatekeeperElodie Lafargue0Jean-Pierre Duneau1Nicolas Buzhinsky2Pamela Ornelas3Alexandre Ortega4Varun Ravishankar5James Sturgis6Ignacio Casuso7Lucie Bergdoll8Turing Centre for Living System, Aix-Marseille Université, U1325 INSERM, DyNaMoLaboratoire d’Ingénierie des Systèmes Macromoléculaires, CNRS - Aix Marseille UniversitéTuring Centre for Living System, Aix-Marseille Université, U1325 INSERM, DyNaMoDepartment of Structural Biology, Max Planck Institute of BiophysicsTuring Centre for Living System, Aix-Marseille Université, U1325 INSERM, DyNaMoLaboratoire d’Ingénierie des Systèmes Macromoléculaires, CNRS - Aix Marseille UniversitéLaboratoire d’Ingénierie des Systèmes Macromoléculaires, CNRS - Aix Marseille UniversitéTuring Centre for Living System, Aix-Marseille Université, U1325 INSERM, DyNaMoLaboratoire d’Ingénierie des Systèmes Macromoléculaires, CNRS - Aix Marseille UniversitéAbstract VDACs, the most abundant proteins in the outer mitochondrial membrane (MOM), are crucial for mitochondrial physiology. VDAC regulate metabolite and ion exchange, modulate calcium homeostasis, and play roles in numerous cellular events such as apoptosis, mitochondrial DNA (mtDNA) release, and different diseases. Mitochondrial function is closely tied to VDAC oligomerization, influencing key processes like mtDNA release and apoptosis, but the molecular drivers of this oligomerization remain unclear. In this study, we investigate the effects of three major MOM lipids on VDAC assemblies using atomic force microscopy and molecular dynamics simulations. Our results show that phosphatidylethanolamine and cholesterol regulate VDAC assembly, with the formation of stable lipid–protein organization of various size and compaction. Deviations from physiological lipid content disrupted native-like VDAC assemblies, highlighting the importance of lipid environment in VDAC organization. These findings underscore how lipid heterogeneity and changes in membranes influence VDAC function.https://doi.org/10.1038/s42003-025-08311-5
spellingShingle Elodie Lafargue
Jean-Pierre Duneau
Nicolas Buzhinsky
Pamela Ornelas
Alexandre Ortega
Varun Ravishankar
James Sturgis
Ignacio Casuso
Lucie Bergdoll
Membrane lipid composition modulates the organization of VDAC1, a mitochondrial gatekeeper
Communications Biology
title Membrane lipid composition modulates the organization of VDAC1, a mitochondrial gatekeeper
title_full Membrane lipid composition modulates the organization of VDAC1, a mitochondrial gatekeeper
title_fullStr Membrane lipid composition modulates the organization of VDAC1, a mitochondrial gatekeeper
title_full_unstemmed Membrane lipid composition modulates the organization of VDAC1, a mitochondrial gatekeeper
title_short Membrane lipid composition modulates the organization of VDAC1, a mitochondrial gatekeeper
title_sort membrane lipid composition modulates the organization of vdac1 a mitochondrial gatekeeper
url https://doi.org/10.1038/s42003-025-08311-5
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