Scale-Dependent Diversity Patterns in Subalpine Grasslands: Homogenization vs. Complexity

This study investigates the scale-dependent alpha and beta diversity patterns in the subalpine grasslands of the Central Balkan Mountains following decades of reduced grazing. We examined two distinct vegetation patches: pure grasslands (N-type) and grasslands mixed with dwarf shrubs (V-type), focus...

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Main Authors: Tsvetelina Terziyska, James Tsakalos, Sándor Bartha, Iva Apostolova, Desislava Sopotlieva
Format: Article
Language:English
Published: MDPI AG 2025-04-01
Series:Land
Subjects:
Online Access:https://www.mdpi.com/2073-445X/14/4/823
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author Tsvetelina Terziyska
James Tsakalos
Sándor Bartha
Iva Apostolova
Desislava Sopotlieva
author_facet Tsvetelina Terziyska
James Tsakalos
Sándor Bartha
Iva Apostolova
Desislava Sopotlieva
author_sort Tsvetelina Terziyska
collection DOAJ
description This study investigates the scale-dependent alpha and beta diversity patterns in the subalpine grasslands of the Central Balkan Mountains following decades of reduced grazing. We examined two distinct vegetation patches: pure grasslands (N-type) and grasslands mixed with dwarf shrubs (V-type), focusing on coarse-scale (among stands) and fine-scale (within stands) heterogeneity. Using micro-quadrat transects and dissimilarity analyses, we assessed the species composition variability and spatial complexity of the two vegetation patches. The results showed that the N-type exhibited higher coarse-scale beta diversity, attributed to fluctuating dominant grass proportions, and lower fine-scale diversity due to competitive exclusion. Conversely, V-type vegetation displayed lower coarse-scale but higher fine-scale diversity, reflecting the heterogeneous spatial matrix created by dwarf-shrub encroachment. Fine-scale spatial complexity, quantified by the compositional diversity of dominants, strongly correlated with species richness and diversity. Two main conclusions emerged: (a) Grazing decline-driven succession toward grass–dwarf shrub mixtures enhanced local diversity while reducing landscape heterogeneity. Conversely, regeneration toward typical dominant grasses diminished local diversity but increased landscape heterogeneity. (b) A balanced fine-scale spatial mixture of dominant species was found to reduce their individual negative impact on other species’ diversity. Effective high-mountain pasture management requires the consideration of scale and local plant co-existence.
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spelling doaj-art-4d5a19fdf7284328bdaf12e2bc411dc42025-08-20T02:28:15ZengMDPI AGLand2073-445X2025-04-0114482310.3390/land14040823Scale-Dependent Diversity Patterns in Subalpine Grasslands: Homogenization vs. ComplexityTsvetelina Terziyska0James Tsakalos1Sándor Bartha2Iva Apostolova3Desislava Sopotlieva4Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Str., 1113 Sofia, BulgariaSchool of Biosciences and Veterinary Medicine, Plant Diversity and Ecosystems Management Unit, University of Camerino, Via Pontoni 5, I-62032 Camerino, MC, ItalyHUN-REN Centre for Ecological Research, Institute of Ecology and Botany, Alkotmány út 2-4, H-2163 Vácrátót, HungaryInstitute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Str., 1113 Sofia, BulgariaInstitute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Str., 1113 Sofia, BulgariaThis study investigates the scale-dependent alpha and beta diversity patterns in the subalpine grasslands of the Central Balkan Mountains following decades of reduced grazing. We examined two distinct vegetation patches: pure grasslands (N-type) and grasslands mixed with dwarf shrubs (V-type), focusing on coarse-scale (among stands) and fine-scale (within stands) heterogeneity. Using micro-quadrat transects and dissimilarity analyses, we assessed the species composition variability and spatial complexity of the two vegetation patches. The results showed that the N-type exhibited higher coarse-scale beta diversity, attributed to fluctuating dominant grass proportions, and lower fine-scale diversity due to competitive exclusion. Conversely, V-type vegetation displayed lower coarse-scale but higher fine-scale diversity, reflecting the heterogeneous spatial matrix created by dwarf-shrub encroachment. Fine-scale spatial complexity, quantified by the compositional diversity of dominants, strongly correlated with species richness and diversity. Two main conclusions emerged: (a) Grazing decline-driven succession toward grass–dwarf shrub mixtures enhanced local diversity while reducing landscape heterogeneity. Conversely, regeneration toward typical dominant grasses diminished local diversity but increased landscape heterogeneity. (b) A balanced fine-scale spatial mixture of dominant species was found to reduce their individual negative impact on other species’ diversity. Effective high-mountain pasture management requires the consideration of scale and local plant co-existence.https://www.mdpi.com/2073-445X/14/4/823alpha diversitybeta diversitydominantsdwarf-shrub encroachmentregenerationsuccession
spellingShingle Tsvetelina Terziyska
James Tsakalos
Sándor Bartha
Iva Apostolova
Desislava Sopotlieva
Scale-Dependent Diversity Patterns in Subalpine Grasslands: Homogenization vs. Complexity
Land
alpha diversity
beta diversity
dominants
dwarf-shrub encroachment
regeneration
succession
title Scale-Dependent Diversity Patterns in Subalpine Grasslands: Homogenization vs. Complexity
title_full Scale-Dependent Diversity Patterns in Subalpine Grasslands: Homogenization vs. Complexity
title_fullStr Scale-Dependent Diversity Patterns in Subalpine Grasslands: Homogenization vs. Complexity
title_full_unstemmed Scale-Dependent Diversity Patterns in Subalpine Grasslands: Homogenization vs. Complexity
title_short Scale-Dependent Diversity Patterns in Subalpine Grasslands: Homogenization vs. Complexity
title_sort scale dependent diversity patterns in subalpine grasslands homogenization vs complexity
topic alpha diversity
beta diversity
dominants
dwarf-shrub encroachment
regeneration
succession
url https://www.mdpi.com/2073-445X/14/4/823
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AT jamestsakalos scaledependentdiversitypatternsinsubalpinegrasslandshomogenizationvscomplexity
AT sandorbartha scaledependentdiversitypatternsinsubalpinegrasslandshomogenizationvscomplexity
AT ivaapostolova scaledependentdiversitypatternsinsubalpinegrasslandshomogenizationvscomplexity
AT desislavasopotlieva scaledependentdiversitypatternsinsubalpinegrasslandshomogenizationvscomplexity