Epilobieae genomes and the evolution of Myrtales

Abstract The order Myrtales, one of the most species-rich lineage within the Superrosidae clade, with the majority of its species distributed across five families: Myrtaceae, Melastomataceae, Lythraceae, Onagraceae, and Combretaceae. Despite the ecological and economic importance of the Myrtales, it...

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Main Authors: Dan Wang, Wenkai Li, Bing Yang, Yujia Shen, Jun He, Derek W. Dunn, Kang Huang, Baoguo Li
Format: Article
Language:English
Published: BMC 2025-07-01
Series:BMC Plant Biology
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Online Access:https://doi.org/10.1186/s12870-025-07003-w
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author Dan Wang
Wenkai Li
Bing Yang
Yujia Shen
Jun He
Derek W. Dunn
Kang Huang
Baoguo Li
author_facet Dan Wang
Wenkai Li
Bing Yang
Yujia Shen
Jun He
Derek W. Dunn
Kang Huang
Baoguo Li
author_sort Dan Wang
collection DOAJ
description Abstract The order Myrtales, one of the most species-rich lineage within the Superrosidae clade, with the majority of its species distributed across five families: Myrtaceae, Melastomataceae, Lythraceae, Onagraceae, and Combretaceae. Despite the ecological and economic importance of the Myrtales, its phylogenetic relationships remain unresolved, with previous studies yielding inconsistent results based on gene fragments and plastid genomes. Genomic data, particularly single-copy/low-copy nuclear genes, provide valuable insights for resolving these phylogenetic relationships. However, phylogenetic studies still lack sufficient clade coverage, particularly for less studied families such as the Onagraceae. Epilobieae is an important tribe of the Onagraceae, characterized by substantial chromosomal number variation and whole genome duplication event (WGD). Nevertheless, the mechanisms of chromosomal evolution remain unresolved. Here, we sequenced and assembled the genomes of two representative species from the Epilobieae, Chamerion angustifolium (formerly Chamaenerion angustifolium) and Epilobium hirsutum, with genome sizes of 636.59 Mb and 400.23 Mb, respectively. Genome evolution analysis revealed two WGD events, during which the chromosome number increased from $$\:\text{n}=9$$ to $$\:\text{n}=18$$ , followed by aneuploid reduction, leading to the diverse chromosomal numbers observed within this tribe. The WGD retained genes are enriched in Environmental information processing pathways, potentially enhancing resistance to biotic and abiotic stresses. These genes also show a preference for multiple exons, which may promote alternative splicing and functional diversification. Additionally, integrating genomic data from 24 Myrtales species, a robust phylogenetic framework based on 994 single-copy/low-copy orthogroups were reconstructed. Our results supported Combretaceae as the sister group to Myrtaceae and Melastomataceae, providing new insights into the evolutionary relationships within the Myrtales.
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spelling doaj-art-4bad49b2ea2c415884fc7e0e66b90e652025-08-20T03:42:27ZengBMCBMC Plant Biology1471-22292025-07-0125111410.1186/s12870-025-07003-wEpilobieae genomes and the evolution of MyrtalesDan Wang0Wenkai Li1Bing Yang2Yujia Shen3Jun He4Derek W. Dunn5Kang Huang6Baoguo Li7Shaanxi Key Laboratory for Animal Conservation, College of Life Sciences, Northwest UniversityShaanxi Key Laboratory for Animal Conservation, College of Life Sciences, Northwest UniversityShaanxi Key Laboratory for Animal Conservation, College of Life Sciences, Northwest UniversityShaanxi Key Laboratory for Animal Conservation, College of Life Sciences, Northwest UniversityCollege of Horticulture, Nanjing Agricultural UniversityShaanxi Key Laboratory for Animal Conservation, College of Life Sciences, Northwest UniversityShaanxi Key Laboratory for Animal Conservation, College of Life Sciences, Northwest UniversityShaanxi Key Laboratory for Animal Conservation, College of Life Sciences, Northwest UniversityAbstract The order Myrtales, one of the most species-rich lineage within the Superrosidae clade, with the majority of its species distributed across five families: Myrtaceae, Melastomataceae, Lythraceae, Onagraceae, and Combretaceae. Despite the ecological and economic importance of the Myrtales, its phylogenetic relationships remain unresolved, with previous studies yielding inconsistent results based on gene fragments and plastid genomes. Genomic data, particularly single-copy/low-copy nuclear genes, provide valuable insights for resolving these phylogenetic relationships. However, phylogenetic studies still lack sufficient clade coverage, particularly for less studied families such as the Onagraceae. Epilobieae is an important tribe of the Onagraceae, characterized by substantial chromosomal number variation and whole genome duplication event (WGD). Nevertheless, the mechanisms of chromosomal evolution remain unresolved. Here, we sequenced and assembled the genomes of two representative species from the Epilobieae, Chamerion angustifolium (formerly Chamaenerion angustifolium) and Epilobium hirsutum, with genome sizes of 636.59 Mb and 400.23 Mb, respectively. Genome evolution analysis revealed two WGD events, during which the chromosome number increased from $$\:\text{n}=9$$ to $$\:\text{n}=18$$ , followed by aneuploid reduction, leading to the diverse chromosomal numbers observed within this tribe. The WGD retained genes are enriched in Environmental information processing pathways, potentially enhancing resistance to biotic and abiotic stresses. These genes also show a preference for multiple exons, which may promote alternative splicing and functional diversification. Additionally, integrating genomic data from 24 Myrtales species, a robust phylogenetic framework based on 994 single-copy/low-copy orthogroups were reconstructed. Our results supported Combretaceae as the sister group to Myrtaceae and Melastomataceae, providing new insights into the evolutionary relationships within the Myrtales.https://doi.org/10.1186/s12870-025-07003-wMyrtalesPhylogenyEpilobieaeChromosome evolutionGene retention
spellingShingle Dan Wang
Wenkai Li
Bing Yang
Yujia Shen
Jun He
Derek W. Dunn
Kang Huang
Baoguo Li
Epilobieae genomes and the evolution of Myrtales
BMC Plant Biology
Myrtales
Phylogeny
Epilobieae
Chromosome evolution
Gene retention
title Epilobieae genomes and the evolution of Myrtales
title_full Epilobieae genomes and the evolution of Myrtales
title_fullStr Epilobieae genomes and the evolution of Myrtales
title_full_unstemmed Epilobieae genomes and the evolution of Myrtales
title_short Epilobieae genomes and the evolution of Myrtales
title_sort epilobieae genomes and the evolution of myrtales
topic Myrtales
Phylogeny
Epilobieae
Chromosome evolution
Gene retention
url https://doi.org/10.1186/s12870-025-07003-w
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