Natural variation in GNP3 determines grain number and grain yield in rice

Abstract Grain number per panicle critically determines rice yield. Although many underlying genes have been reported, yet the molecular mechanisms linking ethylene to panicle development remain unclear. Here, we identify GRAIN NUMBER PER PANICLE 3 (GNP3) as a regulator of GNP through genome-wide as...

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Main Authors: Zhiqi Ma, Yuhang Ming, Jinlong Li, Honglin Li, Haozhen Wang, Xiaoyang Zhu, Yawen Xu, Yong Zhao, Qianfeng Hu, Ruiqi Liu, Yinghua Pan, Danting Li, Wensheng Wang, Jianlong Xu, Xingming Sun, Jinjie Li, Hongliang Zhang, Zichao Li, Zhanying Zhang
Format: Article
Language:English
Published: Nature Portfolio 2025-07-01
Series:Nature Communications
Online Access:https://doi.org/10.1038/s41467-025-61326-8
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author Zhiqi Ma
Yuhang Ming
Jinlong Li
Honglin Li
Haozhen Wang
Xiaoyang Zhu
Yawen Xu
Yong Zhao
Qianfeng Hu
Ruiqi Liu
Yinghua Pan
Danting Li
Wensheng Wang
Jianlong Xu
Xingming Sun
Jinjie Li
Hongliang Zhang
Zichao Li
Zhanying Zhang
author_facet Zhiqi Ma
Yuhang Ming
Jinlong Li
Honglin Li
Haozhen Wang
Xiaoyang Zhu
Yawen Xu
Yong Zhao
Qianfeng Hu
Ruiqi Liu
Yinghua Pan
Danting Li
Wensheng Wang
Jianlong Xu
Xingming Sun
Jinjie Li
Hongliang Zhang
Zichao Li
Zhanying Zhang
author_sort Zhiqi Ma
collection DOAJ
description Abstract Grain number per panicle critically determines rice yield. Although many underlying genes have been reported, yet the molecular mechanisms linking ethylene to panicle development remain unclear. Here, we identify GRAIN NUMBER PER PANICLE 3 (GNP3) as a regulator of GNP through genome-wide association study (GWAS) combined with map-based cloning. GNP3 encodes a MITOGEN-ACTIVATED PROTEIN KINASE KINASE KINASE 22 (OsMKKK22) that phosphorylates S-adenosyl-L-methionine synthetase 1 (SAMS1), triggering its degradation to suppress ethylene biosynthesis. Ethylene overaccumulation in gnp3−1 mutants reduces grain number, while GNP3 overexpression enhances panicle branching and grain yield by lowering ethylene levels. We demonstrate that a natural haplotype GNP3 Hap-Tprevalent in indica subspecies strengthens GNP3-SAMS1 interaction, accelerating SAMS1 degradation and improving grain number. Furthermore, overexpressing GNP3 increases grain yield by approximately 20% in field plot conditions. Our findings unveil a MAPK-ethylene regulatory module and highlight GNP3 Hap-T as a valuable genetic resource for breeding high-yield rice.
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institution Kabale University
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publishDate 2025-07-01
publisher Nature Portfolio
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series Nature Communications
spelling doaj-art-4a14a44b7d7349b8b3ee6bf5d776f8172025-08-20T03:37:37ZengNature PortfolioNature Communications2041-17232025-07-0116111610.1038/s41467-025-61326-8Natural variation in GNP3 determines grain number and grain yield in riceZhiqi Ma0Yuhang Ming1Jinlong Li2Honglin Li3Haozhen Wang4Xiaoyang Zhu5Yawen Xu6Yong Zhao7Qianfeng Hu8Ruiqi Liu9Yinghua Pan10Danting Li11Wensheng Wang12Jianlong Xu13Xingming Sun14Jinjie Li15Hongliang Zhang16Zichao Li17Zhanying Zhang18Frontiers Science Center for Molecular Design Breeding, Key Laboratory of Crop Heterosis and Utilization (MOE), Beijing Key Laboratory of Crop Genetic Improvement, College of Agronomy and Biotechnology, China Agricultural UniversityState Key Laboratory of Plant Environmental Resilience, College of Biological Sciences, China Agricultural UniversityFrontiers Science Center for Molecular Design Breeding, Key Laboratory of Crop Heterosis and Utilization (MOE), Beijing Key Laboratory of Crop Genetic Improvement, College of Agronomy and Biotechnology, China Agricultural UniversityFrontiers Science Center for Molecular Design Breeding, Key Laboratory of Crop Heterosis and Utilization (MOE), Beijing Key Laboratory of Crop Genetic Improvement, College of Agronomy and Biotechnology, China Agricultural UniversityFrontiers Science Center for Molecular Design Breeding, Key Laboratory of Crop Heterosis and Utilization (MOE), Beijing Key Laboratory of Crop Genetic Improvement, College of Agronomy and Biotechnology, China Agricultural UniversityFrontiers Science Center for Molecular Design Breeding, Key Laboratory of Crop Heterosis and Utilization (MOE), Beijing Key Laboratory of Crop Genetic Improvement, College of Agronomy and Biotechnology, China Agricultural UniversityFrontiers Science Center for Molecular Design Breeding, Key Laboratory of Crop Heterosis and Utilization (MOE), Beijing Key Laboratory of Crop Genetic Improvement, College of Agronomy and Biotechnology, China Agricultural UniversityFrontiers Science Center for Molecular Design Breeding, Key Laboratory of Crop Heterosis and Utilization (MOE), Beijing Key Laboratory of Crop Genetic Improvement, College of Agronomy and Biotechnology, China Agricultural UniversityFrontiers Science Center for Molecular Design Breeding, Key Laboratory of Crop Heterosis and Utilization (MOE), Beijing Key Laboratory of Crop Genetic Improvement, College of Agronomy and Biotechnology, China Agricultural UniversityFrontiers Science Center for Molecular Design Breeding, Key Laboratory of Crop Heterosis and Utilization (MOE), Beijing Key Laboratory of Crop Genetic Improvement, College of Agronomy and Biotechnology, China Agricultural UniversityGuangxi Key Laboratory of Rice Genetics and Breeding, Rice Research Institute of Guangxi Academy of Agricultural SciencesGuangxi Key Laboratory of Rice Genetics and Breeding, Rice Research Institute of Guangxi Academy of Agricultural SciencesInstitute of Crop Sciences, Chinese Academy of Agricultural SciencesInstitute of Crop Sciences, Chinese Academy of Agricultural SciencesFrontiers Science Center for Molecular Design Breeding, Key Laboratory of Crop Heterosis and Utilization (MOE), Beijing Key Laboratory of Crop Genetic Improvement, College of Agronomy and Biotechnology, China Agricultural UniversityFrontiers Science Center for Molecular Design Breeding, Key Laboratory of Crop Heterosis and Utilization (MOE), Beijing Key Laboratory of Crop Genetic Improvement, College of Agronomy and Biotechnology, China Agricultural UniversityFrontiers Science Center for Molecular Design Breeding, Key Laboratory of Crop Heterosis and Utilization (MOE), Beijing Key Laboratory of Crop Genetic Improvement, College of Agronomy and Biotechnology, China Agricultural UniversityFrontiers Science Center for Molecular Design Breeding, Key Laboratory of Crop Heterosis and Utilization (MOE), Beijing Key Laboratory of Crop Genetic Improvement, College of Agronomy and Biotechnology, China Agricultural UniversityFrontiers Science Center for Molecular Design Breeding, Key Laboratory of Crop Heterosis and Utilization (MOE), Beijing Key Laboratory of Crop Genetic Improvement, College of Agronomy and Biotechnology, China Agricultural UniversityAbstract Grain number per panicle critically determines rice yield. Although many underlying genes have been reported, yet the molecular mechanisms linking ethylene to panicle development remain unclear. Here, we identify GRAIN NUMBER PER PANICLE 3 (GNP3) as a regulator of GNP through genome-wide association study (GWAS) combined with map-based cloning. GNP3 encodes a MITOGEN-ACTIVATED PROTEIN KINASE KINASE KINASE 22 (OsMKKK22) that phosphorylates S-adenosyl-L-methionine synthetase 1 (SAMS1), triggering its degradation to suppress ethylene biosynthesis. Ethylene overaccumulation in gnp3−1 mutants reduces grain number, while GNP3 overexpression enhances panicle branching and grain yield by lowering ethylene levels. We demonstrate that a natural haplotype GNP3 Hap-Tprevalent in indica subspecies strengthens GNP3-SAMS1 interaction, accelerating SAMS1 degradation and improving grain number. Furthermore, overexpressing GNP3 increases grain yield by approximately 20% in field plot conditions. Our findings unveil a MAPK-ethylene regulatory module and highlight GNP3 Hap-T as a valuable genetic resource for breeding high-yield rice.https://doi.org/10.1038/s41467-025-61326-8
spellingShingle Zhiqi Ma
Yuhang Ming
Jinlong Li
Honglin Li
Haozhen Wang
Xiaoyang Zhu
Yawen Xu
Yong Zhao
Qianfeng Hu
Ruiqi Liu
Yinghua Pan
Danting Li
Wensheng Wang
Jianlong Xu
Xingming Sun
Jinjie Li
Hongliang Zhang
Zichao Li
Zhanying Zhang
Natural variation in GNP3 determines grain number and grain yield in rice
Nature Communications
title Natural variation in GNP3 determines grain number and grain yield in rice
title_full Natural variation in GNP3 determines grain number and grain yield in rice
title_fullStr Natural variation in GNP3 determines grain number and grain yield in rice
title_full_unstemmed Natural variation in GNP3 determines grain number and grain yield in rice
title_short Natural variation in GNP3 determines grain number and grain yield in rice
title_sort natural variation in gnp3 determines grain number and grain yield in rice
url https://doi.org/10.1038/s41467-025-61326-8
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