Formation of complexes at plasmodesmata for potyvirus intercellular movement is mediated by the viral protein P3N-PIPO.

Intercellular transport of viruses through cytoplasmic connections, termed plasmodesmata (PD), is essential for systemic infection in plants by viruses. Previous genetic and ultrastructural data revealed that the potyvirus cyclindrical inclusion (CI) protein is directly involved in cell-to-cell move...

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Main Authors: Taiyun Wei, Changwei Zhang, Jian Hong, Ruyi Xiong, Kristin D Kasschau, Xueping Zhou, James C Carrington, Aiming Wang
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2010-06-01
Series:PLoS Pathogens
Online Access:https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1000962&type=printable
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author Taiyun Wei
Changwei Zhang
Jian Hong
Ruyi Xiong
Kristin D Kasschau
Xueping Zhou
James C Carrington
Aiming Wang
author_facet Taiyun Wei
Changwei Zhang
Jian Hong
Ruyi Xiong
Kristin D Kasschau
Xueping Zhou
James C Carrington
Aiming Wang
author_sort Taiyun Wei
collection DOAJ
description Intercellular transport of viruses through cytoplasmic connections, termed plasmodesmata (PD), is essential for systemic infection in plants by viruses. Previous genetic and ultrastructural data revealed that the potyvirus cyclindrical inclusion (CI) protein is directly involved in cell-to-cell movement, likely through the formation of conical structures anchored to and extended through PD. In this study, we demonstrate that plasmodesmatal localization of CI in N. benthamiana leaf cells is modulated by the recently discovered potyviral protein, P3N-PIPO, in a CI:P3N-PIPO ratio-dependent manner. We show that P3N-PIPO is a PD-located protein that physically interacts with CI in planta. The early secretory pathway, rather than the actomyosin motility system, is required for the delivery of P3N-PIPO and CI to PD. Moreover, CI mutations that disrupt virus cell-to-cell movement compromise PD-localization capacity. These data suggest that the CI and P3N-PIPO complex coordinates the formation of PD-associated structures that facilitate the intercellular movement of potyviruses in infected plants.
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institution DOAJ
issn 1553-7366
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language English
publishDate 2010-06-01
publisher Public Library of Science (PLoS)
record_format Article
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spelling doaj-art-46baadb29b084ec189656c9171f09d8a2025-08-20T03:19:52ZengPublic Library of Science (PLoS)PLoS Pathogens1553-73661553-73742010-06-0166e100096210.1371/journal.ppat.1000962Formation of complexes at plasmodesmata for potyvirus intercellular movement is mediated by the viral protein P3N-PIPO.Taiyun WeiChangwei ZhangJian HongRuyi XiongKristin D KasschauXueping ZhouJames C CarringtonAiming WangIntercellular transport of viruses through cytoplasmic connections, termed plasmodesmata (PD), is essential for systemic infection in plants by viruses. Previous genetic and ultrastructural data revealed that the potyvirus cyclindrical inclusion (CI) protein is directly involved in cell-to-cell movement, likely through the formation of conical structures anchored to and extended through PD. In this study, we demonstrate that plasmodesmatal localization of CI in N. benthamiana leaf cells is modulated by the recently discovered potyviral protein, P3N-PIPO, in a CI:P3N-PIPO ratio-dependent manner. We show that P3N-PIPO is a PD-located protein that physically interacts with CI in planta. The early secretory pathway, rather than the actomyosin motility system, is required for the delivery of P3N-PIPO and CI to PD. Moreover, CI mutations that disrupt virus cell-to-cell movement compromise PD-localization capacity. These data suggest that the CI and P3N-PIPO complex coordinates the formation of PD-associated structures that facilitate the intercellular movement of potyviruses in infected plants.https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1000962&type=printable
spellingShingle Taiyun Wei
Changwei Zhang
Jian Hong
Ruyi Xiong
Kristin D Kasschau
Xueping Zhou
James C Carrington
Aiming Wang
Formation of complexes at plasmodesmata for potyvirus intercellular movement is mediated by the viral protein P3N-PIPO.
PLoS Pathogens
title Formation of complexes at plasmodesmata for potyvirus intercellular movement is mediated by the viral protein P3N-PIPO.
title_full Formation of complexes at plasmodesmata for potyvirus intercellular movement is mediated by the viral protein P3N-PIPO.
title_fullStr Formation of complexes at plasmodesmata for potyvirus intercellular movement is mediated by the viral protein P3N-PIPO.
title_full_unstemmed Formation of complexes at plasmodesmata for potyvirus intercellular movement is mediated by the viral protein P3N-PIPO.
title_short Formation of complexes at plasmodesmata for potyvirus intercellular movement is mediated by the viral protein P3N-PIPO.
title_sort formation of complexes at plasmodesmata for potyvirus intercellular movement is mediated by the viral protein p3n pipo
url https://journals.plos.org/plospathogens/article/file?id=10.1371/journal.ppat.1000962&type=printable
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