CBC Complex Regulates Hyphal Growth, Sclerotial Quantity, and Pathogenicity in the Necrotrophic Fungus <i>Botrytis cinerea</i>

The cap-binding protein complex (CBC), comprising Cbp20 and Cbp80, is crucial for gene expression, yet its role in the notorious crop pathogen <i>Botrytis cinerea</i> remains unclear. Immunoprecipitation coupled with LC-MS/MS demonstrated that BcCbp20 interacts with BcCbp80. Yeast two-hy...

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Main Authors: Yinshan Zhang, Xueting Chen, Guihua Li, Qingming Qin, Mingzhe Zhang, Jianchun Qin
Format: Article
Language:English
Published: MDPI AG 2025-06-01
Series:Journal of Fungi
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Online Access:https://www.mdpi.com/2309-608X/11/6/429
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author Yinshan Zhang
Xueting Chen
Guihua Li
Qingming Qin
Mingzhe Zhang
Jianchun Qin
author_facet Yinshan Zhang
Xueting Chen
Guihua Li
Qingming Qin
Mingzhe Zhang
Jianchun Qin
author_sort Yinshan Zhang
collection DOAJ
description The cap-binding protein complex (CBC), comprising Cbp20 and Cbp80, is crucial for gene expression, yet its role in the notorious crop pathogen <i>Botrytis cinerea</i> remains unclear. Immunoprecipitation coupled with LC-MS/MS demonstrated that BcCbp20 interacts with BcCbp80. Yeast two-hybrid, GST pull-down, and Split-luciferase complementation assays confirmed that the conserved RNA recognition motif (RRM, 54–127 aa) of BcCbp20 and the N-terminal MIF4G domain (1–370 aa, 1–577 aa) of BcCbp80 constitute the core interaction regions. Genetic transformation experiments revealed that <i>BcCBP80</i> exerts a more dominant role than <i>BcCBP20</i> in regulating hyphal morphology, growth rate, conidiophore development, and conidial yield. Furthermore, <i>BcCBP20</i> and <i>BcCBP80</i> differentially regulate sclerotium formation to maintain sclerotial quantity. Based on pathogenicity assays, <i>BcCBP80</i> associated with infection cushion development, with this phenotypic alteration possibly being among the factors correlated with altered pathogenicity. However, the increased sensitivity of Δ<i>Bccbp20</i> to various stress factors may be the primary reason for the diminished pathogenicity. Taken together, these results indicate that <i>BcCBP20</i> and <i>BcCBP80</i> play important roles in multiple aspects of <i>B. cinerea</i> growth, development, stress response, and pathogenicity.
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spelling doaj-art-4575460a4e514311b85ac03548902a412025-08-20T03:27:21ZengMDPI AGJournal of Fungi2309-608X2025-06-0111642910.3390/jof11060429CBC Complex Regulates Hyphal Growth, Sclerotial Quantity, and Pathogenicity in the Necrotrophic Fungus <i>Botrytis cinerea</i>Yinshan Zhang0Xueting Chen1Guihua Li2Qingming Qin3Mingzhe Zhang4Jianchun Qin5College of Plant Science, Jilin University, Changchun 130062, ChinaCollege of Plant Science, Jilin University, Changchun 130062, ChinaCollege of Plant Science, Jilin University, Changchun 130062, ChinaCollege of Plant Science, Jilin University, Changchun 130062, ChinaCollege of Plant Science, Jilin University, Changchun 130062, ChinaCollege of Plant Science, Jilin University, Changchun 130062, ChinaThe cap-binding protein complex (CBC), comprising Cbp20 and Cbp80, is crucial for gene expression, yet its role in the notorious crop pathogen <i>Botrytis cinerea</i> remains unclear. Immunoprecipitation coupled with LC-MS/MS demonstrated that BcCbp20 interacts with BcCbp80. Yeast two-hybrid, GST pull-down, and Split-luciferase complementation assays confirmed that the conserved RNA recognition motif (RRM, 54–127 aa) of BcCbp20 and the N-terminal MIF4G domain (1–370 aa, 1–577 aa) of BcCbp80 constitute the core interaction regions. Genetic transformation experiments revealed that <i>BcCBP80</i> exerts a more dominant role than <i>BcCBP20</i> in regulating hyphal morphology, growth rate, conidiophore development, and conidial yield. Furthermore, <i>BcCBP20</i> and <i>BcCBP80</i> differentially regulate sclerotium formation to maintain sclerotial quantity. Based on pathogenicity assays, <i>BcCBP80</i> associated with infection cushion development, with this phenotypic alteration possibly being among the factors correlated with altered pathogenicity. However, the increased sensitivity of Δ<i>Bccbp20</i> to various stress factors may be the primary reason for the diminished pathogenicity. Taken together, these results indicate that <i>BcCBP20</i> and <i>BcCBP80</i> play important roles in multiple aspects of <i>B. cinerea</i> growth, development, stress response, and pathogenicity.https://www.mdpi.com/2309-608X/11/6/429cap-binding protein complex<i>BcCBP20</i><i>BcCBP80</i>RRMMIF4Ginfection cushion
spellingShingle Yinshan Zhang
Xueting Chen
Guihua Li
Qingming Qin
Mingzhe Zhang
Jianchun Qin
CBC Complex Regulates Hyphal Growth, Sclerotial Quantity, and Pathogenicity in the Necrotrophic Fungus <i>Botrytis cinerea</i>
Journal of Fungi
cap-binding protein complex
<i>BcCBP20</i>
<i>BcCBP80</i>
RRM
MIF4G
infection cushion
title CBC Complex Regulates Hyphal Growth, Sclerotial Quantity, and Pathogenicity in the Necrotrophic Fungus <i>Botrytis cinerea</i>
title_full CBC Complex Regulates Hyphal Growth, Sclerotial Quantity, and Pathogenicity in the Necrotrophic Fungus <i>Botrytis cinerea</i>
title_fullStr CBC Complex Regulates Hyphal Growth, Sclerotial Quantity, and Pathogenicity in the Necrotrophic Fungus <i>Botrytis cinerea</i>
title_full_unstemmed CBC Complex Regulates Hyphal Growth, Sclerotial Quantity, and Pathogenicity in the Necrotrophic Fungus <i>Botrytis cinerea</i>
title_short CBC Complex Regulates Hyphal Growth, Sclerotial Quantity, and Pathogenicity in the Necrotrophic Fungus <i>Botrytis cinerea</i>
title_sort cbc complex regulates hyphal growth sclerotial quantity and pathogenicity in the necrotrophic fungus i botrytis cinerea i
topic cap-binding protein complex
<i>BcCBP20</i>
<i>BcCBP80</i>
RRM
MIF4G
infection cushion
url https://www.mdpi.com/2309-608X/11/6/429
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