Vaccinia virus modulates the redox environment by inhibiting reactive oxygen and nitrogen species with increased activity of endogenous antioxidant enzymes

Abstract The vaccinia virus (VACV) is the most studied and well-characterised member of the Poxviridae family. However, the mechanisms through which it modulates redox homeostasis in host cells remain unclear. Although oxidative stress, which is marked by elevated levels of reactive species, contrib...

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Main Authors: Marília Bueno da Silva Menegatto, Ariane Coelho Ferraz, Rafaela Lameira Souza Lima, Pedro Henrique Guimarães, Oluwashola Samuel Ola-Olu, Pedro Alves Machado-Junior, Wellington Carvalho Malta, Thaís de Fátima Silva Moraes, Frank Silva Bezerra, Breno de Mello Silva, José Carlos de Magalhães, Jordana Grazziela Alves Coelho dos Reis, Flávio Guimarães da Fonseca, Giliane de Souza Trindade, Erna Geessien Kroon, Cintia Lopes de Brito Magalhães
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Language:English
Published: Nature Portfolio 2025-08-01
Series:Scientific Reports
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Online Access:https://doi.org/10.1038/s41598-025-14433-x
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author Marília Bueno da Silva Menegatto
Ariane Coelho Ferraz
Rafaela Lameira Souza Lima
Pedro Henrique Guimarães
Oluwashola Samuel Ola-Olu
Pedro Alves Machado-Junior
Wellington Carvalho Malta
Thaís de Fátima Silva Moraes
Frank Silva Bezerra
Breno de Mello Silva
José Carlos de Magalhães
Jordana Grazziela Alves Coelho dos Reis
Flávio Guimarães da Fonseca
Giliane de Souza Trindade
Erna Geessien Kroon
Cintia Lopes de Brito Magalhães
author_facet Marília Bueno da Silva Menegatto
Ariane Coelho Ferraz
Rafaela Lameira Souza Lima
Pedro Henrique Guimarães
Oluwashola Samuel Ola-Olu
Pedro Alves Machado-Junior
Wellington Carvalho Malta
Thaís de Fátima Silva Moraes
Frank Silva Bezerra
Breno de Mello Silva
José Carlos de Magalhães
Jordana Grazziela Alves Coelho dos Reis
Flávio Guimarães da Fonseca
Giliane de Souza Trindade
Erna Geessien Kroon
Cintia Lopes de Brito Magalhães
author_sort Marília Bueno da Silva Menegatto
collection DOAJ
description Abstract The vaccinia virus (VACV) is the most studied and well-characterised member of the Poxviridae family. However, the mechanisms through which it modulates redox homeostasis in host cells remain unclear. Although oxidative stress, which is marked by elevated levels of reactive species, contributes to the pathogenesis of many viral infections, poxviruses may adopt distinct strategies. VACV has redox effector proteins that are released into the cytosol when the virus penetrates the host cell. This study demonstrates for the first time that VACV infection leads to the activation of the nuclear factor erythroid 2 (Nrf2)/antioxidant response element pathway, a key regulator of cellular antioxidant responses, a mechanism not previously described for any poxvirus. Using BSC-40 cells, we observed that VACV significantly reduced reactive oxygen and nitrogen species levels, downregulated inducible nitric oxide synthase, and enhanced the activity of antioxidant enzymes, such as superoxide dismutase, catalase, and glutathione peroxidase. This antioxidant shift is correlated with increased Nrf2 activity and the upregulation of its downstream targets. This virus-induced antioxidant state may be an immunomodulatory mechanism that facilitates viral replication by dampening host defence. Thus, our findings expand the current understanding of virus–host interactions in poxvirus infections.
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spelling doaj-art-429810cd9f55443fad3d33873f408cbe2025-08-20T03:42:53ZengNature PortfolioScientific Reports2045-23222025-08-0115111210.1038/s41598-025-14433-xVaccinia virus modulates the redox environment by inhibiting reactive oxygen and nitrogen species with increased activity of endogenous antioxidant enzymesMarília Bueno da Silva Menegatto0Ariane Coelho Ferraz1Rafaela Lameira Souza Lima2Pedro Henrique Guimarães3Oluwashola Samuel Ola-Olu4Pedro Alves Machado-Junior5Wellington Carvalho Malta6Thaís de Fátima Silva Moraes7Frank Silva Bezerra8Breno de Mello Silva9José Carlos de Magalhães10Jordana Grazziela Alves Coelho dos Reis11Flávio Guimarães da Fonseca12Giliane de Souza Trindade13Erna Geessien Kroon14Cintia Lopes de Brito Magalhães15Programa de Pós Graduação em Ciências Biológicas, Núcleo de Pesquisas em Ciências Biológicas, NUPEB, Universidade Federal de Ouro PretoPrograma de Pós Graduação em Ciências Biológicas, Núcleo de Pesquisas em Ciências Biológicas, NUPEB, Universidade Federal de Ouro PretoPrograma de Pós Graduação em Ciências Biológicas, Núcleo de Pesquisas em Ciências Biológicas, NUPEB, Universidade Federal de Ouro PretoPrograma de Pós Graduação em Ciências Biológicas, Núcleo de Pesquisas em Ciências Biológicas, NUPEB, Universidade Federal de Ouro PretoPrograma de Pós Graduação em Biotecnologia, Núcleo de Pesquisas em Ciências Biológicas, NUPEB, Universidade Federal de Ouro PretoPrograma de Pós Graduação em Ciências Biológicas, Núcleo de Pesquisas em Ciências Biológicas, NUPEB, Universidade Federal de Ouro PretoPrograma de Pós Graduação em Biotecnologia, Núcleo de Pesquisas em Ciências Biológicas, NUPEB, Universidade Federal de Ouro PretoPrograma de Pós Graduação em Microbiologia, Universidade Federal de Minas GeraisPrograma de Pós Graduação em Ciências Biológicas, Núcleo de Pesquisas em Ciências Biológicas, NUPEB, Universidade Federal de Ouro PretoPrograma de Pós Graduação em Ciências Biológicas, Núcleo de Pesquisas em Ciências Biológicas, NUPEB, Universidade Federal de Ouro PretoPrograma de Pós Graduação em Biotecnologia, Universidade Federal de São João del-ReiPrograma de Pós Graduação em Microbiologia, Universidade Federal de Minas GeraisPrograma de Pós Graduação em Microbiologia, Universidade Federal de Minas GeraisPrograma de Pós Graduação em Microbiologia, Universidade Federal de Minas GeraisPrograma de Pós Graduação em Microbiologia, Universidade Federal de Minas GeraisPrograma de Pós Graduação em Ciências Biológicas, Núcleo de Pesquisas em Ciências Biológicas, NUPEB, Universidade Federal de Ouro PretoAbstract The vaccinia virus (VACV) is the most studied and well-characterised member of the Poxviridae family. However, the mechanisms through which it modulates redox homeostasis in host cells remain unclear. Although oxidative stress, which is marked by elevated levels of reactive species, contributes to the pathogenesis of many viral infections, poxviruses may adopt distinct strategies. VACV has redox effector proteins that are released into the cytosol when the virus penetrates the host cell. This study demonstrates for the first time that VACV infection leads to the activation of the nuclear factor erythroid 2 (Nrf2)/antioxidant response element pathway, a key regulator of cellular antioxidant responses, a mechanism not previously described for any poxvirus. Using BSC-40 cells, we observed that VACV significantly reduced reactive oxygen and nitrogen species levels, downregulated inducible nitric oxide synthase, and enhanced the activity of antioxidant enzymes, such as superoxide dismutase, catalase, and glutathione peroxidase. This antioxidant shift is correlated with increased Nrf2 activity and the upregulation of its downstream targets. This virus-induced antioxidant state may be an immunomodulatory mechanism that facilitates viral replication by dampening host defence. Thus, our findings expand the current understanding of virus–host interactions in poxvirus infections.https://doi.org/10.1038/s41598-025-14433-xVaccinia virusPoxvirusesAntioxidantRedox homeostasis
spellingShingle Marília Bueno da Silva Menegatto
Ariane Coelho Ferraz
Rafaela Lameira Souza Lima
Pedro Henrique Guimarães
Oluwashola Samuel Ola-Olu
Pedro Alves Machado-Junior
Wellington Carvalho Malta
Thaís de Fátima Silva Moraes
Frank Silva Bezerra
Breno de Mello Silva
José Carlos de Magalhães
Jordana Grazziela Alves Coelho dos Reis
Flávio Guimarães da Fonseca
Giliane de Souza Trindade
Erna Geessien Kroon
Cintia Lopes de Brito Magalhães
Vaccinia virus modulates the redox environment by inhibiting reactive oxygen and nitrogen species with increased activity of endogenous antioxidant enzymes
Scientific Reports
Vaccinia virus
Poxviruses
Antioxidant
Redox homeostasis
title Vaccinia virus modulates the redox environment by inhibiting reactive oxygen and nitrogen species with increased activity of endogenous antioxidant enzymes
title_full Vaccinia virus modulates the redox environment by inhibiting reactive oxygen and nitrogen species with increased activity of endogenous antioxidant enzymes
title_fullStr Vaccinia virus modulates the redox environment by inhibiting reactive oxygen and nitrogen species with increased activity of endogenous antioxidant enzymes
title_full_unstemmed Vaccinia virus modulates the redox environment by inhibiting reactive oxygen and nitrogen species with increased activity of endogenous antioxidant enzymes
title_short Vaccinia virus modulates the redox environment by inhibiting reactive oxygen and nitrogen species with increased activity of endogenous antioxidant enzymes
title_sort vaccinia virus modulates the redox environment by inhibiting reactive oxygen and nitrogen species with increased activity of endogenous antioxidant enzymes
topic Vaccinia virus
Poxviruses
Antioxidant
Redox homeostasis
url https://doi.org/10.1038/s41598-025-14433-x
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