Chloroplast Z-ring dynamics is governed by conserved core regions of evolutionarily divergent FtsZs

The chloroplast FtsZ ring (Z ring) is assembled by two distinct FtsZ proteins, FtsZ2 and FtsZ1 (referred to as FtsZA and FtsZB in red algae). FtsZ2 confers stability to the Z ring, while FtsZ1 enhances its dynamics. Enhanced Z-ring dynamics is essential for Z-ring remodeling, which drives chloroplas...

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Main Authors: Lingyan Cao, Katie J. Porter, Wenbin Du, Emily J. Tallerday, Moyang Liu, Wanqi Liang, Katherine W. Osteryoung, Cheng Chen
Format: Article
Language:English
Published: Frontiers Media S.A. 2025-07-01
Series:Frontiers in Plant Science
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Online Access:https://www.frontiersin.org/articles/10.3389/fpls.2025.1622675/full
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author Lingyan Cao
Lingyan Cao
Lingyan Cao
Katie J. Porter
Wenbin Du
Emily J. Tallerday
Moyang Liu
Wanqi Liang
Katherine W. Osteryoung
Cheng Chen
Cheng Chen
Cheng Chen
author_facet Lingyan Cao
Lingyan Cao
Lingyan Cao
Katie J. Porter
Wenbin Du
Emily J. Tallerday
Moyang Liu
Wanqi Liang
Katherine W. Osteryoung
Cheng Chen
Cheng Chen
Cheng Chen
author_sort Lingyan Cao
collection DOAJ
description The chloroplast FtsZ ring (Z ring) is assembled by two distinct FtsZ proteins, FtsZ2 and FtsZ1 (referred to as FtsZA and FtsZB in red algae). FtsZ2 confers stability to the Z ring, while FtsZ1 enhances its dynamics. Enhanced Z-ring dynamics is essential for Z-ring remodeling, which drives chloroplast constriction and division. However, the mechanisms underlying the distinct dynamic properties of the two FtsZs remain unclear. Here, we report that the conserved core regions are primarily responsible for the distinct dynamic properties observed in both plant and red algal FtsZs. We demonstrate that the conserved core region of FtsZ1 enhances the dynamics of FtsZ2 within coassembled filaments. Likewise, we show that the conserved core region of red algal FtsZB promotes the dynamics of coassembled FtsZA rings. Our findings provide evidence that the evolution of a second FtsZ protein represents a general mechanism to enhance the dynamics of the chloroplast Z ring.
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institution Kabale University
issn 1664-462X
language English
publishDate 2025-07-01
publisher Frontiers Media S.A.
record_format Article
series Frontiers in Plant Science
spelling doaj-art-41d2c179e0fd4f358599dfdd1702cdfa2025-08-20T03:56:09ZengFrontiers Media S.A.Frontiers in Plant Science1664-462X2025-07-011610.3389/fpls.2025.16226751622675Chloroplast Z-ring dynamics is governed by conserved core regions of evolutionarily divergent FtsZsLingyan Cao0Lingyan Cao1Lingyan Cao2Katie J. Porter3Wenbin Du4Emily J. Tallerday5Moyang Liu6Wanqi Liang7Katherine W. Osteryoung8Cheng Chen9Cheng Chen10Cheng Chen11Joint International Research Laboratory of Metabolic and Developmental Sciences, State Key Laboratory of Hybrid Rice, School of Life Sciences and Biotechnology, Shanghai Jiao Tong University, Shanghai, ChinaDepartment of Plant Biology, Michigan State University, East Lansing, MI, United StatesDepartment of Plant Sciences, School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, ChinaDepartment of Plant Biology, Michigan State University, East Lansing, MI, United StatesDepartment of Plant Sciences, School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, ChinaDepartment of Plant Biology, Michigan State University, East Lansing, MI, United StatesDepartment of Plant Sciences, School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, ChinaJoint International Research Laboratory of Metabolic and Developmental Sciences, State Key Laboratory of Hybrid Rice, School of Life Sciences and Biotechnology, Shanghai Jiao Tong University, Shanghai, ChinaDepartment of Plant Biology, Michigan State University, East Lansing, MI, United StatesDepartment of Plant Sciences, School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, ChinaShanghai Collaborative Innovation Center of Agri-Seeds, Shanghai Jiao Tong University, Shanghai, ChinaJoint Center for Single Cell Biology, Shanghai Jiao Tong University, Shanghai, ChinaThe chloroplast FtsZ ring (Z ring) is assembled by two distinct FtsZ proteins, FtsZ2 and FtsZ1 (referred to as FtsZA and FtsZB in red algae). FtsZ2 confers stability to the Z ring, while FtsZ1 enhances its dynamics. Enhanced Z-ring dynamics is essential for Z-ring remodeling, which drives chloroplast constriction and division. However, the mechanisms underlying the distinct dynamic properties of the two FtsZs remain unclear. Here, we report that the conserved core regions are primarily responsible for the distinct dynamic properties observed in both plant and red algal FtsZs. We demonstrate that the conserved core region of FtsZ1 enhances the dynamics of FtsZ2 within coassembled filaments. Likewise, we show that the conserved core region of red algal FtsZB promotes the dynamics of coassembled FtsZA rings. Our findings provide evidence that the evolution of a second FtsZ protein represents a general mechanism to enhance the dynamics of the chloroplast Z ring.https://www.frontiersin.org/articles/10.3389/fpls.2025.1622675/fullchloroplastFtsZ-ringdynamicsdivisionevolution
spellingShingle Lingyan Cao
Lingyan Cao
Lingyan Cao
Katie J. Porter
Wenbin Du
Emily J. Tallerday
Moyang Liu
Wanqi Liang
Katherine W. Osteryoung
Cheng Chen
Cheng Chen
Cheng Chen
Chloroplast Z-ring dynamics is governed by conserved core regions of evolutionarily divergent FtsZs
Frontiers in Plant Science
chloroplast
FtsZ-ring
dynamics
division
evolution
title Chloroplast Z-ring dynamics is governed by conserved core regions of evolutionarily divergent FtsZs
title_full Chloroplast Z-ring dynamics is governed by conserved core regions of evolutionarily divergent FtsZs
title_fullStr Chloroplast Z-ring dynamics is governed by conserved core regions of evolutionarily divergent FtsZs
title_full_unstemmed Chloroplast Z-ring dynamics is governed by conserved core regions of evolutionarily divergent FtsZs
title_short Chloroplast Z-ring dynamics is governed by conserved core regions of evolutionarily divergent FtsZs
title_sort chloroplast z ring dynamics is governed by conserved core regions of evolutionarily divergent ftszs
topic chloroplast
FtsZ-ring
dynamics
division
evolution
url https://www.frontiersin.org/articles/10.3389/fpls.2025.1622675/full
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