Melatonin Mitigates Cd-Induced Growth Repression and RNA m<sup>6</sup>A Hypermethylation by Triggering MMR-Mediated DNA Damage Response

Melatonin (MT) has been found to mitigate cadmium (Cd) toxicity with negligible environmental risks. It remains poorly understood as to how MT mitigates Cd-induced growth repression and regulates RNA m<sup>6</sup>A methylation. We aimed to elucidate the effect of MT on growth repression...

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Main Authors: Zihan Tang, Hetong Wang, Xianpeng Wang, Richard A. Ludlow, Zhouli Liu, Min Zhang, Qijiang Cao, Wan Liu, Qiang Zhao
Format: Article
Language:English
Published: MDPI AG 2025-05-01
Series:Plants
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Online Access:https://www.mdpi.com/2223-7747/14/9/1398
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author Zihan Tang
Hetong Wang
Xianpeng Wang
Richard A. Ludlow
Zhouli Liu
Min Zhang
Qijiang Cao
Wan Liu
Qiang Zhao
author_facet Zihan Tang
Hetong Wang
Xianpeng Wang
Richard A. Ludlow
Zhouli Liu
Min Zhang
Qijiang Cao
Wan Liu
Qiang Zhao
author_sort Zihan Tang
collection DOAJ
description Melatonin (MT) has been found to mitigate cadmium (Cd) toxicity with negligible environmental risks. It remains poorly understood as to how MT mitigates Cd-induced growth repression and regulates RNA m<sup>6</sup>A methylation. We aimed to elucidate the effect of MT on growth repression and RNA m<sup>6</sup>A methylation in Arabidopsis (<i>Arabidopsis thaliana</i>) exposed to Cd stress. MT mitigated, on average, 13.96% and 8.42% of growth repression resulting from Cd and mismatch repair (MMR) deficiency. The ameliorative effect on Cd stress was reduced by 70.56% and 34.23% in <i>msh2</i> and <i>msh6</i> mutants, respectively. With distinct dose–effect relationships, m<sup>6</sup>A hypermethylation responded to Cd stress rather than Cu stress, which was further elevated in MMR-deficient seedlings. MT reduced m<sup>6</sup>A levels by 22.98% even without stress induction, whereas the depressed m<sup>6</sup>A levels in MMR-deficient seedlings, greatly exceeding those in the WT. The “writer” and “eraser” gene expression responsible for m<sup>6</sup>A methylation was reduced with the concentration of stresses due to MT, but <i>VIR</i> and <i>ALKBH9B</i> no longer responded to Cd stress in <i>msh2</i> and <i>msh6</i>. Despite the remarkable repression, MMR gene expression was regularly promoted by MT under Cd and Cu stress. Our study provides novel insights into the molecular mechanisms underlying the restorative effects of MT on growth repression and m<sup>6</sup>A methylation regulation, which shed light on Cd phytoremediation.
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spelling doaj-art-40f608fecba94adcad0f8a366ab7ae192025-08-20T03:52:57ZengMDPI AGPlants2223-77472025-05-01149139810.3390/plants14091398Melatonin Mitigates Cd-Induced Growth Repression and RNA m<sup>6</sup>A Hypermethylation by Triggering MMR-Mediated DNA Damage ResponseZihan Tang0Hetong Wang1Xianpeng Wang2Richard A. Ludlow3Zhouli Liu4Min Zhang5Qijiang Cao6Wan Liu7Qiang Zhao8Liaoning Key Laboratory of Urban Integrated Pest Management and Ecological Security, College of Life Science and Bioengineering, Shenyang University, Shenyang 110044, ChinaLiaoning Key Laboratory of Urban Integrated Pest Management and Ecological Security, College of Life Science and Bioengineering, Shenyang University, Shenyang 110044, ChinaLiaoning Key Laboratory of Urban Integrated Pest Management and Ecological Security, College of Life Science and Bioengineering, Shenyang University, Shenyang 110044, ChinaSchool of Biosciences, Cardiff University, Sir Martin Evans Building, Museum Avenue, Cardiff CF10 3AX, UKLiaoning Key Laboratory of Urban Integrated Pest Management and Ecological Security, College of Life Science and Bioengineering, Shenyang University, Shenyang 110044, ChinaLiaoning Key Laboratory of Urban Integrated Pest Management and Ecological Security, College of Life Science and Bioengineering, Shenyang University, Shenyang 110044, ChinaLiaoning Key Laboratory of Urban Integrated Pest Management and Ecological Security, College of Life Science and Bioengineering, Shenyang University, Shenyang 110044, ChinaCollege of Agriculture and Biological Sciences, Dehong Normal University, Mangshi 678400, ChinaCollege of Agriculture, Heilongjiang Bayi Agricultural University, Daqing 163000, ChinaMelatonin (MT) has been found to mitigate cadmium (Cd) toxicity with negligible environmental risks. It remains poorly understood as to how MT mitigates Cd-induced growth repression and regulates RNA m<sup>6</sup>A methylation. We aimed to elucidate the effect of MT on growth repression and RNA m<sup>6</sup>A methylation in Arabidopsis (<i>Arabidopsis thaliana</i>) exposed to Cd stress. MT mitigated, on average, 13.96% and 8.42% of growth repression resulting from Cd and mismatch repair (MMR) deficiency. The ameliorative effect on Cd stress was reduced by 70.56% and 34.23% in <i>msh2</i> and <i>msh6</i> mutants, respectively. With distinct dose–effect relationships, m<sup>6</sup>A hypermethylation responded to Cd stress rather than Cu stress, which was further elevated in MMR-deficient seedlings. MT reduced m<sup>6</sup>A levels by 22.98% even without stress induction, whereas the depressed m<sup>6</sup>A levels in MMR-deficient seedlings, greatly exceeding those in the WT. The “writer” and “eraser” gene expression responsible for m<sup>6</sup>A methylation was reduced with the concentration of stresses due to MT, but <i>VIR</i> and <i>ALKBH9B</i> no longer responded to Cd stress in <i>msh2</i> and <i>msh6</i>. Despite the remarkable repression, MMR gene expression was regularly promoted by MT under Cd and Cu stress. Our study provides novel insights into the molecular mechanisms underlying the restorative effects of MT on growth repression and m<sup>6</sup>A methylation regulation, which shed light on Cd phytoremediation.https://www.mdpi.com/2223-7747/14/9/1398Cd toxicologyRNA m<sup>6</sup>A methylationDNA repaircell cycleArabidopsis
spellingShingle Zihan Tang
Hetong Wang
Xianpeng Wang
Richard A. Ludlow
Zhouli Liu
Min Zhang
Qijiang Cao
Wan Liu
Qiang Zhao
Melatonin Mitigates Cd-Induced Growth Repression and RNA m<sup>6</sup>A Hypermethylation by Triggering MMR-Mediated DNA Damage Response
Plants
Cd toxicology
RNA m<sup>6</sup>A methylation
DNA repair
cell cycle
Arabidopsis
title Melatonin Mitigates Cd-Induced Growth Repression and RNA m<sup>6</sup>A Hypermethylation by Triggering MMR-Mediated DNA Damage Response
title_full Melatonin Mitigates Cd-Induced Growth Repression and RNA m<sup>6</sup>A Hypermethylation by Triggering MMR-Mediated DNA Damage Response
title_fullStr Melatonin Mitigates Cd-Induced Growth Repression and RNA m<sup>6</sup>A Hypermethylation by Triggering MMR-Mediated DNA Damage Response
title_full_unstemmed Melatonin Mitigates Cd-Induced Growth Repression and RNA m<sup>6</sup>A Hypermethylation by Triggering MMR-Mediated DNA Damage Response
title_short Melatonin Mitigates Cd-Induced Growth Repression and RNA m<sup>6</sup>A Hypermethylation by Triggering MMR-Mediated DNA Damage Response
title_sort melatonin mitigates cd induced growth repression and rna m sup 6 sup a hypermethylation by triggering mmr mediated dna damage response
topic Cd toxicology
RNA m<sup>6</sup>A methylation
DNA repair
cell cycle
Arabidopsis
url https://www.mdpi.com/2223-7747/14/9/1398
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