Enhanced flux potential analysis links changes in enzyme expression to metabolic flux

Abstract Algorithms that constrain metabolic network models with enzyme levels to predict metabolic activity assume that changes in enzyme levels are indicative of flux variations. However, metabolic flux can also be regulated by other mechanisms such as allostery and mass action. To systematically...

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Main Authors: Xuhang Li, Albertha J M Walhout, L Safak Yilmaz
Format: Article
Language:English
Published: Springer Nature 2025-02-01
Series:Molecular Systems Biology
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Online Access:https://doi.org/10.1038/s44320-025-00090-9
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author Xuhang Li
Albertha J M Walhout
L Safak Yilmaz
author_facet Xuhang Li
Albertha J M Walhout
L Safak Yilmaz
author_sort Xuhang Li
collection DOAJ
description Abstract Algorithms that constrain metabolic network models with enzyme levels to predict metabolic activity assume that changes in enzyme levels are indicative of flux variations. However, metabolic flux can also be regulated by other mechanisms such as allostery and mass action. To systematically explore the relationship between fluctuations in enzyme expression and flux, we combine available yeast proteomic and fluxomic data to reveal that flux changes can be best predicted from changes in enzyme levels of pathways, rather than the whole network or only cognate reactions. We implement this principle in an ‘enhanced flux potential analysis’ (eFPA) algorithm that integrates enzyme expression data with metabolic network architecture to predict relative flux levels of reactions including those regulated by other mechanisms. Applied to human data, eFPA consistently predicts tissue metabolic function using either proteomic or transcriptomic data. Additionally, eFPA efficiently handles data sparsity and noisiness, generating robust flux predictions with single-cell gene expression data. Our approach outperforms alternatives by striking an optimal balance, evaluating enzyme expression at pathway level, rather than either single-reaction or whole-network levels.
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spelling doaj-art-390b1c207ca541bfa15cbab1465f41c12025-08-20T03:03:20ZengSpringer NatureMolecular Systems Biology1744-42922025-02-0121441344510.1038/s44320-025-00090-9Enhanced flux potential analysis links changes in enzyme expression to metabolic fluxXuhang Li0Albertha J M Walhout1L Safak Yilmaz2Department of Systems Biology, University of Massachusetts Chan Medical SchoolDepartment of Systems Biology, University of Massachusetts Chan Medical SchoolDepartment of Systems Biology, University of Massachusetts Chan Medical SchoolAbstract Algorithms that constrain metabolic network models with enzyme levels to predict metabolic activity assume that changes in enzyme levels are indicative of flux variations. However, metabolic flux can also be regulated by other mechanisms such as allostery and mass action. To systematically explore the relationship between fluctuations in enzyme expression and flux, we combine available yeast proteomic and fluxomic data to reveal that flux changes can be best predicted from changes in enzyme levels of pathways, rather than the whole network or only cognate reactions. We implement this principle in an ‘enhanced flux potential analysis’ (eFPA) algorithm that integrates enzyme expression data with metabolic network architecture to predict relative flux levels of reactions including those regulated by other mechanisms. Applied to human data, eFPA consistently predicts tissue metabolic function using either proteomic or transcriptomic data. Additionally, eFPA efficiently handles data sparsity and noisiness, generating robust flux predictions with single-cell gene expression data. Our approach outperforms alternatives by striking an optimal balance, evaluating enzyme expression at pathway level, rather than either single-reaction or whole-network levels.https://doi.org/10.1038/s44320-025-00090-9Enzyme ExpressionMetabolic Network ModelMetabolic FluxFlux Potential AnalysisSingle-cell Data
spellingShingle Xuhang Li
Albertha J M Walhout
L Safak Yilmaz
Enhanced flux potential analysis links changes in enzyme expression to metabolic flux
Molecular Systems Biology
Enzyme Expression
Metabolic Network Model
Metabolic Flux
Flux Potential Analysis
Single-cell Data
title Enhanced flux potential analysis links changes in enzyme expression to metabolic flux
title_full Enhanced flux potential analysis links changes in enzyme expression to metabolic flux
title_fullStr Enhanced flux potential analysis links changes in enzyme expression to metabolic flux
title_full_unstemmed Enhanced flux potential analysis links changes in enzyme expression to metabolic flux
title_short Enhanced flux potential analysis links changes in enzyme expression to metabolic flux
title_sort enhanced flux potential analysis links changes in enzyme expression to metabolic flux
topic Enzyme Expression
Metabolic Network Model
Metabolic Flux
Flux Potential Analysis
Single-cell Data
url https://doi.org/10.1038/s44320-025-00090-9
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AT lsafakyilmaz enhancedfluxpotentialanalysislinkschangesinenzymeexpressiontometabolicflux