A Comprehensive Analysis of Short Specific Tissue (SST) Proteins, a New Group of Proteins from PF10950 That May Give Rise to Cyclopeptide Alkaloids

Proteins of the PF10950 family feature the DUF2775 domain of unknown function. The most studied are specific tissue (ST) proteins with tandem repeats, which are putative precursors of cyclopeptide alkaloids. Here, we study uncharacterised short ST (SST) proteins with the DUFF2775 domain by analysing...

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Main Authors: Lucía Albornos, Paula Iriondo, Silvia Rodríguez-Marcos, Patricia Farelo, Guillermo Sobrino-Mengual, Luz María Muñoz-Centeno, Ignacio Martín, Berta Dopico
Format: Article
Language:English
Published: MDPI AG 2025-04-01
Series:Plants
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Online Access:https://www.mdpi.com/2223-7747/14/7/1117
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author Lucía Albornos
Paula Iriondo
Silvia Rodríguez-Marcos
Patricia Farelo
Guillermo Sobrino-Mengual
Luz María Muñoz-Centeno
Ignacio Martín
Berta Dopico
author_facet Lucía Albornos
Paula Iriondo
Silvia Rodríguez-Marcos
Patricia Farelo
Guillermo Sobrino-Mengual
Luz María Muñoz-Centeno
Ignacio Martín
Berta Dopico
author_sort Lucía Albornos
collection DOAJ
description Proteins of the PF10950 family feature the DUF2775 domain of unknown function. The most studied are specific tissue (ST) proteins with tandem repeats, which are putative precursors of cyclopeptide alkaloids. Here, we study uncharacterised short ST (SST) proteins with the DUFF2775 domain by analysing 194 sequences from 120 species of 39 taxonomic families in silico. SST proteins have a signal peptide and their size and several other characteristics depend on their individual taxonomic family. Sequence analyses revealed that SST proteins contain two well-conserved regions, one resembling the ST repeat, which could constitute the core of cyclopeptide alkaloids. We studied the unique <i>SST1</i> gene of <i>Arabidopsis thaliana</i>, which is adjacent to and co-expressed with a gene encoding a protein with a BURP domain, associated with cyclopeptide production. The empirical analysis indicated that the <i>SST1</i> promoter is mainly activated in the roots, where most of the transcripts accumulate, and that the SST1 protein accumulates in the root vascular cambium. At the cellular level, SST fused to GFP appears in vesicles that co-localise with the endoplasmic reticulum and the vacuole. Thus, SSTs are a new type of PF10950 protein found in core eudicots with two conserved regions that could be involved in root biology.
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spelling doaj-art-35a006704de544799cdcbdcba9efaa582025-08-20T03:03:27ZengMDPI AGPlants2223-77472025-04-01147111710.3390/plants14071117A Comprehensive Analysis of Short Specific Tissue (SST) Proteins, a New Group of Proteins from PF10950 That May Give Rise to Cyclopeptide AlkaloidsLucía Albornos0Paula Iriondo1Silvia Rodríguez-Marcos2Patricia Farelo3Guillermo Sobrino-Mengual4Luz María Muñoz-Centeno5Ignacio Martín6Berta Dopico7Department of Botany and Plant Physiology, University of Salamanca, Campus Miguel de Unamuno, 37007 Salamanca, SpainDepartment of Botany and Plant Physiology, University of Salamanca, Campus Miguel de Unamuno, 37007 Salamanca, SpainDepartment of Botany and Plant Physiology, University of Salamanca, Campus Miguel de Unamuno, 37007 Salamanca, SpainDepartment of Botany and Plant Physiology, University of Salamanca, Campus Miguel de Unamuno, 37007 Salamanca, SpainDepartment of Botany and Plant Physiology, University of Salamanca, Campus Miguel de Unamuno, 37007 Salamanca, SpainDepartment of Botany and Plant Physiology, University of Salamanca, Campus Miguel de Unamuno, 37007 Salamanca, SpainDepartment of Botany and Plant Physiology, University of Salamanca, Campus Miguel de Unamuno, 37007 Salamanca, SpainDepartment of Botany and Plant Physiology, University of Salamanca, Campus Miguel de Unamuno, 37007 Salamanca, SpainProteins of the PF10950 family feature the DUF2775 domain of unknown function. The most studied are specific tissue (ST) proteins with tandem repeats, which are putative precursors of cyclopeptide alkaloids. Here, we study uncharacterised short ST (SST) proteins with the DUFF2775 domain by analysing 194 sequences from 120 species of 39 taxonomic families in silico. SST proteins have a signal peptide and their size and several other characteristics depend on their individual taxonomic family. Sequence analyses revealed that SST proteins contain two well-conserved regions, one resembling the ST repeat, which could constitute the core of cyclopeptide alkaloids. We studied the unique <i>SST1</i> gene of <i>Arabidopsis thaliana</i>, which is adjacent to and co-expressed with a gene encoding a protein with a BURP domain, associated with cyclopeptide production. The empirical analysis indicated that the <i>SST1</i> promoter is mainly activated in the roots, where most of the transcripts accumulate, and that the SST1 protein accumulates in the root vascular cambium. At the cellular level, SST fused to GFP appears in vesicles that co-localise with the endoplasmic reticulum and the vacuole. Thus, SSTs are a new type of PF10950 protein found in core eudicots with two conserved regions that could be involved in root biology.https://www.mdpi.com/2223-7747/14/7/1117DUF2775 domainSST proteinST proteinrootcyclopeptides alkaloidsPF10950
spellingShingle Lucía Albornos
Paula Iriondo
Silvia Rodríguez-Marcos
Patricia Farelo
Guillermo Sobrino-Mengual
Luz María Muñoz-Centeno
Ignacio Martín
Berta Dopico
A Comprehensive Analysis of Short Specific Tissue (SST) Proteins, a New Group of Proteins from PF10950 That May Give Rise to Cyclopeptide Alkaloids
Plants
DUF2775 domain
SST protein
ST protein
root
cyclopeptides alkaloids
PF10950
title A Comprehensive Analysis of Short Specific Tissue (SST) Proteins, a New Group of Proteins from PF10950 That May Give Rise to Cyclopeptide Alkaloids
title_full A Comprehensive Analysis of Short Specific Tissue (SST) Proteins, a New Group of Proteins from PF10950 That May Give Rise to Cyclopeptide Alkaloids
title_fullStr A Comprehensive Analysis of Short Specific Tissue (SST) Proteins, a New Group of Proteins from PF10950 That May Give Rise to Cyclopeptide Alkaloids
title_full_unstemmed A Comprehensive Analysis of Short Specific Tissue (SST) Proteins, a New Group of Proteins from PF10950 That May Give Rise to Cyclopeptide Alkaloids
title_short A Comprehensive Analysis of Short Specific Tissue (SST) Proteins, a New Group of Proteins from PF10950 That May Give Rise to Cyclopeptide Alkaloids
title_sort comprehensive analysis of short specific tissue sst proteins a new group of proteins from pf10950 that may give rise to cyclopeptide alkaloids
topic DUF2775 domain
SST protein
ST protein
root
cyclopeptides alkaloids
PF10950
url https://www.mdpi.com/2223-7747/14/7/1117
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