FM dye photo-oxidation as a tool for monitoring membrane recycling in inner hair cells.

Styryl (FM) dyes have been used for more than two decades to investigate exo- and endocytosis in conventional synapses. However, they are difficult to use in the inner hair cells of the auditory pathway (IHCs), as FM dyes appear to penetrate through mechanotransducer channels into the cytosol of IHC...

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Main Authors: Dirk Kamin, Natalia H Revelo, Silvio O Rizzoli
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2014-01-01
Series:PLoS ONE
Online Access:https://doi.org/10.1371/journal.pone.0088353
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author Dirk Kamin
Natalia H Revelo
Silvio O Rizzoli
author_facet Dirk Kamin
Natalia H Revelo
Silvio O Rizzoli
author_sort Dirk Kamin
collection DOAJ
description Styryl (FM) dyes have been used for more than two decades to investigate exo- and endocytosis in conventional synapses. However, they are difficult to use in the inner hair cells of the auditory pathway (IHCs), as FM dyes appear to penetrate through mechanotransducer channels into the cytosol of IHCs, masking endocytotic uptake. To solve this problem we applied to IHCs the FM dye photo-oxidation technique, which renders the dyes into electron microscopy markers. Photo-oxidation allowed the unambiguous identification of labeled organelles, despite the presence of FM dye in the cytosol. This enabled us to describe the morphologies of several organelles that take up membrane in IHCs, both at rest and during stimulation. At rest, endosome-like organelles were detected in the region of the cuticular plate. Larger tubulo-cisternal organelles dominated the top and nuclear regions. Finally, the basal region, where the IHC active zones are located, contained few labeled organelles. Stimulation increased significantly membrane trafficking in the basal region, inducing the appearance of labeled vesicles and cistern-like organelles. The latter were replaced by small, synaptic-like vesicles during recovery after stimulation. In contrast, no changes in membrane trafficking were induced by stimulation in the cuticular plate region or in the top and nuclear regions. We conclude that synaptic vesicle recycling takes place mostly in the basal region of the IHCs. Other organelles participate in abundant constitutive membrane trafficking throughout the rest of the IHC volume.
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spelling doaj-art-319f7a591f6648549852231273c8b6b62025-08-20T03:46:13ZengPublic Library of Science (PLoS)PLoS ONE1932-62032014-01-0192e8835310.1371/journal.pone.0088353FM dye photo-oxidation as a tool for monitoring membrane recycling in inner hair cells.Dirk KaminNatalia H ReveloSilvio O RizzoliStyryl (FM) dyes have been used for more than two decades to investigate exo- and endocytosis in conventional synapses. However, they are difficult to use in the inner hair cells of the auditory pathway (IHCs), as FM dyes appear to penetrate through mechanotransducer channels into the cytosol of IHCs, masking endocytotic uptake. To solve this problem we applied to IHCs the FM dye photo-oxidation technique, which renders the dyes into electron microscopy markers. Photo-oxidation allowed the unambiguous identification of labeled organelles, despite the presence of FM dye in the cytosol. This enabled us to describe the morphologies of several organelles that take up membrane in IHCs, both at rest and during stimulation. At rest, endosome-like organelles were detected in the region of the cuticular plate. Larger tubulo-cisternal organelles dominated the top and nuclear regions. Finally, the basal region, where the IHC active zones are located, contained few labeled organelles. Stimulation increased significantly membrane trafficking in the basal region, inducing the appearance of labeled vesicles and cistern-like organelles. The latter were replaced by small, synaptic-like vesicles during recovery after stimulation. In contrast, no changes in membrane trafficking were induced by stimulation in the cuticular plate region or in the top and nuclear regions. We conclude that synaptic vesicle recycling takes place mostly in the basal region of the IHCs. Other organelles participate in abundant constitutive membrane trafficking throughout the rest of the IHC volume.https://doi.org/10.1371/journal.pone.0088353
spellingShingle Dirk Kamin
Natalia H Revelo
Silvio O Rizzoli
FM dye photo-oxidation as a tool for monitoring membrane recycling in inner hair cells.
PLoS ONE
title FM dye photo-oxidation as a tool for monitoring membrane recycling in inner hair cells.
title_full FM dye photo-oxidation as a tool for monitoring membrane recycling in inner hair cells.
title_fullStr FM dye photo-oxidation as a tool for monitoring membrane recycling in inner hair cells.
title_full_unstemmed FM dye photo-oxidation as a tool for monitoring membrane recycling in inner hair cells.
title_short FM dye photo-oxidation as a tool for monitoring membrane recycling in inner hair cells.
title_sort fm dye photo oxidation as a tool for monitoring membrane recycling in inner hair cells
url https://doi.org/10.1371/journal.pone.0088353
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AT silvioorizzoli fmdyephotooxidationasatoolformonitoringmembranerecyclingininnerhaircells