HIV-1 replication through hHR23A-mediated interaction of Vpr with 26S proteasome.

HIV-1 Vpr is a virion-associated protein. Its activities link to viral pathogenesis and disease progression of HIV-infected patients. In vitro, Vpr moderately activates HIV-1 replication in proliferating T cells, but it is required for efficient viral infection and replication in vivo in non-dividin...

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Main Authors: Ge Li, Robert T Elder, Larisa Dubrovsky, Dong Liang, Tatiana Pushkarsky, Karen Chiu, Tao Fan, Josephine Sire, Michael Bukrinsky, Richard Y Zhao
Format: Article
Language:English
Published: Public Library of Science (PLoS) 2010-06-01
Series:PLoS ONE
Online Access:https://journals.plos.org/plosone/article/file?id=10.1371/journal.pone.0011371&type=printable
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author Ge Li
Robert T Elder
Larisa Dubrovsky
Dong Liang
Tatiana Pushkarsky
Karen Chiu
Tao Fan
Josephine Sire
Michael Bukrinsky
Richard Y Zhao
author_facet Ge Li
Robert T Elder
Larisa Dubrovsky
Dong Liang
Tatiana Pushkarsky
Karen Chiu
Tao Fan
Josephine Sire
Michael Bukrinsky
Richard Y Zhao
author_sort Ge Li
collection DOAJ
description HIV-1 Vpr is a virion-associated protein. Its activities link to viral pathogenesis and disease progression of HIV-infected patients. In vitro, Vpr moderately activates HIV-1 replication in proliferating T cells, but it is required for efficient viral infection and replication in vivo in non-dividing cells such as macrophages. How exactly Vpr contributes to viral replication remains elusive. We show here that Vpr stimulates HIV-1 replication at least in part through its interaction with hHR23A, a protein that binds to 19S subunit of the 26S proteasome and shuttles ubiquitinated proteins to the proteasome for degradation. The Vpr-proteasome interaction was initially discovered in fission yeast, where Vpr was shown to associate with Mts4 and Mts2, two 19S-associated proteins. The interaction of Vpr with the 19S subunit of the proteasome was further confirmed in mammalian cells where Vpr associates with the mammalian orthologues of fission yeast Mts4 and S5a. Consistently, depletion of hHR23A interrupts interaction of Vpr with proteasome in mammalian cells. Furthermore, Vpr promotes hHR23A-mediated protein-ubiquitination, and down-regulation of hHR23A using RNAi significantly reduced viral replication in non-proliferating MAGI-CCR5 cells and primary macrophages. These findings suggest that Vpr-proteasome interaction might counteract certain host restriction factor(s) to stimulate viral replication in non-dividing cells.
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spelling doaj-art-313e82aa97d542b7932285f675fac46c2025-08-20T02:01:56ZengPublic Library of Science (PLoS)PLoS ONE1932-62032010-06-0156e1137110.1371/journal.pone.0011371HIV-1 replication through hHR23A-mediated interaction of Vpr with 26S proteasome.Ge LiRobert T ElderLarisa DubrovskyDong LiangTatiana PushkarskyKaren ChiuTao FanJosephine SireMichael BukrinskyRichard Y ZhaoHIV-1 Vpr is a virion-associated protein. Its activities link to viral pathogenesis and disease progression of HIV-infected patients. In vitro, Vpr moderately activates HIV-1 replication in proliferating T cells, but it is required for efficient viral infection and replication in vivo in non-dividing cells such as macrophages. How exactly Vpr contributes to viral replication remains elusive. We show here that Vpr stimulates HIV-1 replication at least in part through its interaction with hHR23A, a protein that binds to 19S subunit of the 26S proteasome and shuttles ubiquitinated proteins to the proteasome for degradation. The Vpr-proteasome interaction was initially discovered in fission yeast, where Vpr was shown to associate with Mts4 and Mts2, two 19S-associated proteins. The interaction of Vpr with the 19S subunit of the proteasome was further confirmed in mammalian cells where Vpr associates with the mammalian orthologues of fission yeast Mts4 and S5a. Consistently, depletion of hHR23A interrupts interaction of Vpr with proteasome in mammalian cells. Furthermore, Vpr promotes hHR23A-mediated protein-ubiquitination, and down-regulation of hHR23A using RNAi significantly reduced viral replication in non-proliferating MAGI-CCR5 cells and primary macrophages. These findings suggest that Vpr-proteasome interaction might counteract certain host restriction factor(s) to stimulate viral replication in non-dividing cells.https://journals.plos.org/plosone/article/file?id=10.1371/journal.pone.0011371&type=printable
spellingShingle Ge Li
Robert T Elder
Larisa Dubrovsky
Dong Liang
Tatiana Pushkarsky
Karen Chiu
Tao Fan
Josephine Sire
Michael Bukrinsky
Richard Y Zhao
HIV-1 replication through hHR23A-mediated interaction of Vpr with 26S proteasome.
PLoS ONE
title HIV-1 replication through hHR23A-mediated interaction of Vpr with 26S proteasome.
title_full HIV-1 replication through hHR23A-mediated interaction of Vpr with 26S proteasome.
title_fullStr HIV-1 replication through hHR23A-mediated interaction of Vpr with 26S proteasome.
title_full_unstemmed HIV-1 replication through hHR23A-mediated interaction of Vpr with 26S proteasome.
title_short HIV-1 replication through hHR23A-mediated interaction of Vpr with 26S proteasome.
title_sort hiv 1 replication through hhr23a mediated interaction of vpr with 26s proteasome
url https://journals.plos.org/plosone/article/file?id=10.1371/journal.pone.0011371&type=printable
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