Defining neuronal responses to the neurotropic parasite Toxoplasma gondii

ABSTRACT A select group of pathogens infects neurons in the brain. Prior dogma held that neurons were “defenseless” against infecting microbes, but many studies suggest that neurons can mount anti-microbial defenses. However, a knowledge gap in understanding how neurons respond in vitro and in vivo...

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Main Authors: Hannah J. Johnson, Joshua A. Kochanowsky, Sambamurthy Chandrasekaran, Christopher A. Hunter, Daniel P. Beiting, Anita A. Koshy
Format: Article
Language:English
Published: American Society for Microbiology 2025-06-01
Series:mSphere
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Online Access:https://journals.asm.org/doi/10.1128/msphere.00216-25
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author Hannah J. Johnson
Joshua A. Kochanowsky
Sambamurthy Chandrasekaran
Christopher A. Hunter
Daniel P. Beiting
Anita A. Koshy
author_facet Hannah J. Johnson
Joshua A. Kochanowsky
Sambamurthy Chandrasekaran
Christopher A. Hunter
Daniel P. Beiting
Anita A. Koshy
author_sort Hannah J. Johnson
collection DOAJ
description ABSTRACT A select group of pathogens infects neurons in the brain. Prior dogma held that neurons were “defenseless” against infecting microbes, but many studies suggest that neurons can mount anti-microbial defenses. However, a knowledge gap in understanding how neurons respond in vitro and in vivo to different classes of microorganisms remains. To address this gap, we compared a transcriptional data set derived from primary neuron cultures (PNCs) infected with the neurotropic intracellular parasite Toxoplasma gondii with a data set derived from neurons injected with T. gondii protein in vivo. These curated responses were then compared to the transcriptional responses of PNCs infected with the single-stranded RNA viruses, West Nile virus or Zika virus. These analyses highlighted a conserved response to infection associated with chemokines (Cxcl10, Ccl2) and cytokines (interferon signaling). However, T. gondii had diminished IFN-α signaling in vitro compared to the viral data sets and was uniquely associated with a decrease in neuron-specific genes (Snap25, Slc17a7, Prkcg). These data underscore that neurons participate in infection-induced neuroinflammation and illustrate that neurons possess both pathogen-specific and pathogen-conserved responses.IMPORTANCEThough neurons are commonly the target of pathogens that infect the central nervous system (CNS), few data sets assess the neuronal response to infection. This paucity of data is likely because neurons are perceived to have diminished immune capabilities. However, to understand the role of neurons in neuroinflammation and their immune capabilities, their responses must be investigated. Here, we analyzed publicly accessible, neuron-specific data sets to compare neuron responses to a eukaryotic pathogen vs two Orthoflaviviruses. A better understanding of neuron responses to different infections will allow us to develop methods for inhibiting pathways that lead to neuron dysfunction, enhancing those that limit pathogen survival, and mitigating infection-induced damage to the CNS.
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spelling doaj-art-30eb6dcd7fd54a88a93d9aaa20d5158c2025-08-20T03:23:51ZengAmerican Society for MicrobiologymSphere2379-50422025-06-0110610.1128/msphere.00216-25Defining neuronal responses to the neurotropic parasite Toxoplasma gondiiHannah J. Johnson0Joshua A. Kochanowsky1Sambamurthy Chandrasekaran2Christopher A. Hunter3Daniel P. Beiting4Anita A. Koshy5Neuroscience Graduate Interdisciplinary Program, University of Arizona, Tucson, Arizona, USABIO5 Institute, University of Arizona, Tucson, Arizona, USABIO5 Institute, University of Arizona, Tucson, Arizona, USADepartment of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, USADepartment of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, USANeuroscience Graduate Interdisciplinary Program, University of Arizona, Tucson, Arizona, USAABSTRACT A select group of pathogens infects neurons in the brain. Prior dogma held that neurons were “defenseless” against infecting microbes, but many studies suggest that neurons can mount anti-microbial defenses. However, a knowledge gap in understanding how neurons respond in vitro and in vivo to different classes of microorganisms remains. To address this gap, we compared a transcriptional data set derived from primary neuron cultures (PNCs) infected with the neurotropic intracellular parasite Toxoplasma gondii with a data set derived from neurons injected with T. gondii protein in vivo. These curated responses were then compared to the transcriptional responses of PNCs infected with the single-stranded RNA viruses, West Nile virus or Zika virus. These analyses highlighted a conserved response to infection associated with chemokines (Cxcl10, Ccl2) and cytokines (interferon signaling). However, T. gondii had diminished IFN-α signaling in vitro compared to the viral data sets and was uniquely associated with a decrease in neuron-specific genes (Snap25, Slc17a7, Prkcg). These data underscore that neurons participate in infection-induced neuroinflammation and illustrate that neurons possess both pathogen-specific and pathogen-conserved responses.IMPORTANCEThough neurons are commonly the target of pathogens that infect the central nervous system (CNS), few data sets assess the neuronal response to infection. This paucity of data is likely because neurons are perceived to have diminished immune capabilities. However, to understand the role of neurons in neuroinflammation and their immune capabilities, their responses must be investigated. Here, we analyzed publicly accessible, neuron-specific data sets to compare neuron responses to a eukaryotic pathogen vs two Orthoflaviviruses. A better understanding of neuron responses to different infections will allow us to develop methods for inhibiting pathways that lead to neuron dysfunction, enhancing those that limit pathogen survival, and mitigating infection-induced damage to the CNS.https://journals.asm.org/doi/10.1128/msphere.00216-25Toxoplasma gondiiT. gondiineuronsRNA-seqtranscriptomicshost response
spellingShingle Hannah J. Johnson
Joshua A. Kochanowsky
Sambamurthy Chandrasekaran
Christopher A. Hunter
Daniel P. Beiting
Anita A. Koshy
Defining neuronal responses to the neurotropic parasite Toxoplasma gondii
mSphere
Toxoplasma gondii
T. gondii
neurons
RNA-seq
transcriptomics
host response
title Defining neuronal responses to the neurotropic parasite Toxoplasma gondii
title_full Defining neuronal responses to the neurotropic parasite Toxoplasma gondii
title_fullStr Defining neuronal responses to the neurotropic parasite Toxoplasma gondii
title_full_unstemmed Defining neuronal responses to the neurotropic parasite Toxoplasma gondii
title_short Defining neuronal responses to the neurotropic parasite Toxoplasma gondii
title_sort defining neuronal responses to the neurotropic parasite toxoplasma gondii
topic Toxoplasma gondii
T. gondii
neurons
RNA-seq
transcriptomics
host response
url https://journals.asm.org/doi/10.1128/msphere.00216-25
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