Salmonella exploits host- and bacterial-derived β-alanine for replication inside host macrophages

Salmonella is a major foodborne pathogen that can effectively replicate inside host macrophages to establish life-threatening systemic infections. Salmonella must utilize diverse nutrients for growth in nutrient-poor macrophages, but which nutrients are required for intracellular Salmonella growth i...

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Main Authors: Shuai Ma, Bin Yang, Yuyang Sun, Xinyue Wang, Houliang Guo, Ruiying Liu, Ting Ye, Chenbo Kang, Jingnan Chen, Lingyan Jiang
Format: Article
Language:English
Published: eLife Sciences Publications Ltd 2025-06-01
Series:eLife
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Online Access:https://elifesciences.org/articles/103714
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author Shuai Ma
Bin Yang
Yuyang Sun
Xinyue Wang
Houliang Guo
Ruiying Liu
Ting Ye
Chenbo Kang
Jingnan Chen
Lingyan Jiang
author_facet Shuai Ma
Bin Yang
Yuyang Sun
Xinyue Wang
Houliang Guo
Ruiying Liu
Ting Ye
Chenbo Kang
Jingnan Chen
Lingyan Jiang
author_sort Shuai Ma
collection DOAJ
description Salmonella is a major foodborne pathogen that can effectively replicate inside host macrophages to establish life-threatening systemic infections. Salmonella must utilize diverse nutrients for growth in nutrient-poor macrophages, but which nutrients are required for intracellular Salmonella growth is largely unknown. Here, we found that either acquisition from the host or de novo synthesis of a nonprotein amino acid, β-alanine, is critical for Salmonella replication inside macrophages. The concentration of β-alanine is decreased in Salmonella-infected macrophages, while the addition of exogenous β-alanine enhances Salmonella replication in macrophages, suggesting that Salmonella can uptake host-derived β-alanine for intracellular growth. Moreover, the expression of panD, the rate-limiting gene required for β-alanine synthesis in Salmonella, is upregulated when Salmonella enters macrophages. Mutation of panD impaired Salmonella replication in macrophages and colonization in the mouse liver and spleen, indicating that de novo synthesis of β-alanine is essential for intracellular Salmonella growth and systemic infection. Additionally, we revealed that β-alanine influences Salmonella intracellular replication and in vivo virulence partially by increasing expression of the zinc transporter genes znuABC, which in turn facilitates the uptake of the essential micronutrient zinc by Salmonella. Taken together, these findings highlight the important role of β-alanine in the intracellular replication and virulence of Salmonella, and panD is a promising target for controlling systemic Salmonella infection.
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spelling doaj-art-2f9f2e01978947aba3fa86901a52f8be2025-08-20T03:31:16ZengeLife Sciences Publications LtdeLife2050-084X2025-06-011310.7554/eLife.103714Salmonella exploits host- and bacterial-derived β-alanine for replication inside host macrophagesShuai Ma0https://orcid.org/0009-0006-8348-0805Bin Yang1Yuyang Sun2Xinyue Wang3Houliang Guo4Ruiying Liu5Ting Ye6Chenbo Kang7Jingnan Chen8Lingyan Jiang9https://orcid.org/0000-0001-8580-606XNational Key Laboratory of Intelligent Tracking and Forecasting for Infectious Diseases, TEDA Institute of Biological Sciences and Biotechnology, Nankai University, Tianjin, ChinaNational Key Laboratory of Intelligent Tracking and Forecasting for Infectious Diseases, TEDA Institute of Biological Sciences and Biotechnology, Nankai University, Tianjin, ChinaNational Key Laboratory of Intelligent Tracking and Forecasting for Infectious Diseases, TEDA Institute of Biological Sciences and Biotechnology, Nankai University, Tianjin, ChinaNational Key Laboratory of Intelligent Tracking and Forecasting for Infectious Diseases, TEDA Institute of Biological Sciences and Biotechnology, Nankai University, Tianjin, ChinaNational Key Laboratory of Intelligent Tracking and Forecasting for Infectious Diseases, TEDA Institute of Biological Sciences and Biotechnology, Nankai University, Tianjin, ChinaNational Key Laboratory of Intelligent Tracking and Forecasting for Infectious Diseases, TEDA Institute of Biological Sciences and Biotechnology, Nankai University, Tianjin, ChinaNational Key Laboratory of Intelligent Tracking and Forecasting for Infectious Diseases, TEDA Institute of Biological Sciences and Biotechnology, Nankai University, Tianjin, ChinaNational Key Laboratory of Intelligent Tracking and Forecasting for Infectious Diseases, TEDA Institute of Biological Sciences and Biotechnology, Nankai University, Tianjin, ChinaNational Key Laboratory of Intelligent Tracking and Forecasting for Infectious Diseases, TEDA Institute of Biological Sciences and Biotechnology, Nankai University, Tianjin, ChinaNational Key Laboratory of Intelligent Tracking and Forecasting for Infectious Diseases, TEDA Institute of Biological Sciences and Biotechnology, Nankai University, Tianjin, ChinaSalmonella is a major foodborne pathogen that can effectively replicate inside host macrophages to establish life-threatening systemic infections. Salmonella must utilize diverse nutrients for growth in nutrient-poor macrophages, but which nutrients are required for intracellular Salmonella growth is largely unknown. Here, we found that either acquisition from the host or de novo synthesis of a nonprotein amino acid, β-alanine, is critical for Salmonella replication inside macrophages. The concentration of β-alanine is decreased in Salmonella-infected macrophages, while the addition of exogenous β-alanine enhances Salmonella replication in macrophages, suggesting that Salmonella can uptake host-derived β-alanine for intracellular growth. Moreover, the expression of panD, the rate-limiting gene required for β-alanine synthesis in Salmonella, is upregulated when Salmonella enters macrophages. Mutation of panD impaired Salmonella replication in macrophages and colonization in the mouse liver and spleen, indicating that de novo synthesis of β-alanine is essential for intracellular Salmonella growth and systemic infection. Additionally, we revealed that β-alanine influences Salmonella intracellular replication and in vivo virulence partially by increasing expression of the zinc transporter genes znuABC, which in turn facilitates the uptake of the essential micronutrient zinc by Salmonella. Taken together, these findings highlight the important role of β-alanine in the intracellular replication and virulence of Salmonella, and panD is a promising target for controlling systemic Salmonella infection.https://elifesciences.org/articles/103714Salmonellareplication in macrophagevirulenceβ-alaninezinc uptake
spellingShingle Shuai Ma
Bin Yang
Yuyang Sun
Xinyue Wang
Houliang Guo
Ruiying Liu
Ting Ye
Chenbo Kang
Jingnan Chen
Lingyan Jiang
Salmonella exploits host- and bacterial-derived β-alanine for replication inside host macrophages
eLife
Salmonella
replication in macrophage
virulence
β-alanine
zinc uptake
title Salmonella exploits host- and bacterial-derived β-alanine for replication inside host macrophages
title_full Salmonella exploits host- and bacterial-derived β-alanine for replication inside host macrophages
title_fullStr Salmonella exploits host- and bacterial-derived β-alanine for replication inside host macrophages
title_full_unstemmed Salmonella exploits host- and bacterial-derived β-alanine for replication inside host macrophages
title_short Salmonella exploits host- and bacterial-derived β-alanine for replication inside host macrophages
title_sort salmonella exploits host and bacterial derived β alanine for replication inside host macrophages
topic Salmonella
replication in macrophage
virulence
β-alanine
zinc uptake
url https://elifesciences.org/articles/103714
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